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Volume I (1961) - Flora of New Zealand Indigenous Tracheophyta - Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledons
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Aciphylla squarrosa J.R.Forst. & G.Forst.

A. squarrosa J. R. et G. Forst. Char. Gen. Pl. 1776, 136, t. 68.

Type locality: Totaranui, Queen Charlotte Sound. Type: BM.

Us. large plants up to c. 1 m. tall, forming dense tussocks. Lvs ∞, 3-pinnate, spreading, subglaucous; margins and midribs crenulate-serrulate; occ. plants may have some lvs 2-pinnate. Sheaths erect, whitish, fleshy except membr. margins, up to 15 × 5 cm.; stipules with petiolate bases c. 2 × 1 cm., unequal, 3-foliolate, outer segs up to 20 cm., inner to 6 cm., terminal to 25 cm. long, pungent; petiolar or ligular portion sts up to c. 4cm. long. Petioles up to c. 15 cm. × 1-2 mm., internodes up to c. 5 cm. long. Primary pinnae us. 1 pair, again pinnate, internodes c. 3 cm. long; secondary pinnae us. 1 pair, again pinnate, sts bearing accessory pinnules; final segs up to 25 cm. × 4 mm., concavo-convex, with long pungent points. Stems of male plants ridged, up to ± 8 dm. × 2 cm. including infl. up to c. 4 dm. long. Bracts. ∞; sheaths pale, membr., ridged, up to 4 × 1 cm., tapering to apex. stipules unequal, up to 3 cm. × 1 mm. Lamina 3-foliolate; lateral pair of lflts up to c. 9 cm. long; terminal lflt to 20-25 cm. × 3 mm., stiff, pungent, becoming strongly reflexed. Umbels on stout peduncles up to c. 6-10 cm. long, us. with lateral umbellules near base as well as at apex, up to c. 4 cm. diam. Umbellules crowded, forming heads ± 2 cm. diam; rays ∞, ± 10 mm. long. Stems of female plants up to ± 1 m. tall including infl. up to c. 6 dm. tall. Bract-sheaths thin, ribbed; lower c. 5 × 2 cm. stipules unequal, 4-8-(15) cm. long. Lamina of lower sheaths pinnate, with occ. accessory pinnules; petiole flat, ± 10 mm. long, internodes to 2 cm. long; pinnae 2 pairs; terminal lflt up to 15 cm. long, at length strongly reflexed. Umbels c. 8, on stout peduncles ± 5 cm. long bearing lateral umbellules near base; involucral bracts linear. Umbellules ± 10, c. 15 mm. diam., on short rays. Petals pale yellow. Fr. c. 6-8 mm. long; mericarps 4-3-winged. Vittae 2-3 per furrow, 4-6 commissural.

DIST.: N., S. Coastal to montane from Mount Hikurangi to c. lat 41º 30'.

A somewhat polymorphic sp., especially as regards size. Fruiting specimens may be found with lvs not > 15 cm. long, with stems not > 4 dm. tall including infl. ± 2 dm. long; sts the whole blade of the bracts may be ± reflexed. The lvs may be quite glaucous or greyish green, the stipules variously developed.

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