Volume V (2000) - Flora of New Zealand Gramineae
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Festuca L.

Festuca L., 1753

Type species: F. ovina L.

Perennial caespitose, stoloniferous, or rhizomatous. Leaf-sheath split to base or margins fused, apical auricles small. Ligule ciliolate. Leaf-blade convolute, or folded; usually persistent; pungent in F. ultramafica; variable in TS. Culms unbranched; nodes glabrous. Inflorescence an open or contracted panicle of 2-many-flowered laterally compressed, pedicelled spikelets disarticulating above the glumes. Glumes unequal or subequal, persistent, ± keeled and ornamented, rarely exceeding lemmas. Lemma lanceolate-acute or awned, rounded on back or slightly carinate, membranous to thinly coriaceous, 5-7-nerved, awn terminal. Palea bidentate, keels scabrid. Flowers ⚥, usually chasmogamous. Lodicules 2, unequally lobed, often hair-tipped. Stamens 3. Ovary glabrous or hispid hairy at apex. Caryopsis dorsiconvex, ventrally sulcate, free; embryo small; hilum long, linear. Fig. 6.

Key

1
Branching intravaginal
2
Branching extravaginal
9
2
Leaf-blades antrorsely scabrid
3
Leaf-blades smooth
5
3
Lemma awnless or mucronate; spikelets 4-7 mm
Lemma evidently awned; spikelets 8-20 mm
4
4
Panicle with erect or lax branches not conspicuously borne above glaucous brown leaves
Panicle with wide angled branches conspicuously borne above glaucous blue leaves
4a
Rachis prickle-toothed on margins and abundantly minutely so elsewhere
Rachis prickle-toothed on angular margins
5
Panicle ± open; glumes not including lemmas
6
Panicle contracted; glumes including lemmas
8
6
Lemma margins conspicuously hair-fringed
Lemma margins exceedingly finely prickle-toothed
7
7
Leaf-blades persisting on sheath
7a
Leaf-blades thick; stout stature; culm internodes glabrous; spikelets 14-20 mm
Leaf-blades thin; slender stature; culm internodes glabrous or prickle-toothed; spikelets 10-12 mm
Leaf-blades disarticulating at collar
8
Awns 6-13 mm, ≥ lemma; anthers 3.75-4.25 mm; chasmogamous
Awns 1.5-2.5 mm, « lemma; anthers 0.6-0.8 mm; cleistogamous
9
Lemma glabrous
10
Lemma prickle-toothed and/or hairy
12
10
Panicle branches wide-angled
Panicle branches spreading-erect, occasionally otherwise
11
11
Leaf-blades weakly hexagonal or terete, secund; awns 0-1.5 mm
Leaf-blades conduplicate, pungent; awns 1.5-4.5 mm
12
Glume apex conspicuously ciliate
Glume apex pointed
13
13
Anthers 0.5-1.0 mm; flowers cleistogamous
Anthers 2-3 mm; flowers chasmogamous
13a
Habit rhizomatous
Habit caespitose

c. 450 spp. worldwide - "A large and variable genus for which no overall treatment is available..." (Clayton and Renvoize 1986 op. cit.). Endemic spp. 9, indigenous sp. 1; naturalised spp. 3.

The genus was revised for N.Z. indigenous species by Connor, H. E. N.Z. J. Bot. 36: 329-367 (1998); Howarth, W. O. J. Linn. Soc. Lond. 48: 57-77 (1928) - incorrectly cited as vol. 68 in Connor 1998 (op. cit.) - and Saint-Yves, A. Candollea 4: 293-307 (1931) had earlier treated endemic and naturalised taxa. Variation in leaf-blade anatomy for F. matthewsii and F. novae-zelandiae is detailed in Connor, H. E., N.Z. J. Sci. 3: 468-509 (1960). Experimental hybrids between F. matthewsii and F. novae-zelandiae are fertile [Connor, H. E. N.Z. J. Bot. 6: 293-308 (1968)]; none is known in the wild.

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