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Volume IV (1988) - Flora of New Zealand Naturalised Pteridophytes, Gymnosperms, Dicotyledons
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Glossary

GLOSSARY

GLOSSARY

abaxial: facing away from the axis; used of lower lf surface.

abscissing: falling off; often used of floral parts.

acaulescent, acaulous: lacking a distinct stem.

accrescent: enlarging toward maturity, as of calyx after flowering.

accumbent: lying closely face to face; a radicle lying against the edges of a cotyledon.

acentric: not located at the centre.

achene: a small, dry, indehiscent, 1-seeded fr. with thin pericarp.

acicle: needle-shaped process; hence acicular.

acropetally: developing in a longitudinal plane from a lower to a more apical level.

acroscopic: of a pinna or lobe pointing toward the apex.

actinomorphic: with the parts radially symmetrical.

acuminate: tapering to a fine point, the sides ± concave.

acute: sharply pointed.

adaxial: facing towards the axis; used of upper lf surface.

adnate: united to an organ of a different kind, as sepals to petals.

adventitious: arising irregularly or in an unusual position, as roots from stems.

aerial: arising in the air, as roots from branches.

aestivation: the arrangement of perianth parts in the bud; see imbricate, valvate.

agamospecies: microspp. reproducing by apomixis.

allopolyploid: a polyploid derived from a hybrid between 2 or more taxa.

alluvial: of deposits of earth, sand, etc. left by water.

alpine: the area above tree limit.

alternate: spaced singly along an axis, not in opposite pairs or whorls; usually of lvs.

alveolae: deep, close pits; hence alveolate.

amphibious: living on both land and in water.

amphitropous: of ovules, retaining an essentially straight axis but turned through 90° on and fused with funiculus.

amplexicaul: clasping the stem.

ampliate: enlarged.

analgesics: drugs used to alleviate pain.

anastomosing: joining up to form loops or a network; usually of veins.

anatropous: of ovules, turned through 180° and with funiculus fused along one side and micropyle close to placental surface.

androgynophore: a stalk bearing both stamens and pistil above the point of perianth attachment.

andromonoecious: with ♂ and ⚥ fls on the same plant.

androphore: a stalk bearing the stamens.

anemophilous: pollinated via wind.

angustate: of lvs narrowed at the base.

angustiseptate: of a flattened 2-locular capsule (or silique) with the septum across the narrow diameter.

annual: completing its life cycle within 1 year.

annulus: a ring; particularly the ring of thick-walled cells in a fern sporangium forming part of the dehiscence mechanism.

anterior: on the side away from the axis and so appearing in front.

antheridia: in Cryptogams, the organs producing ♂ gametes.

anthocarp: an achene enclosed by the persistent base of the calyx.

anthocyanin: pigment producing blue or purple coloration.

antipetalous: positioned opposite the petals; usually of stamens.

antisepalous: positioned opposite the sepals; usually of stamens.

antrorse: directed forward or upward; usually of hairs.

apetalous: without petals.

apetiolate: without a petiole; hence apetiolulate, of leaflets.

apex: the tip or free end of an organ or part; hence apical, apically.

apiculus: a short, slender, ± flexible point; hence apiculate.

apocarpous: having free carpels.

apomict: plant producing viable seed without fertilization; hence apomictic.

apophysis: the exposed, expanded, distal part of the ovuliferous scale in ♀ conifer cones.

arborescent: approaching the size and habit of a tree.

archegonia: in Cryptogams, the organs producing ♀ gametes.

arcuate: curved or arched, ± strongly.

areole: a small delimited area, as the space between anastomosing veins; hence areolate.

aril: an appendage to a seed, usually an outgrowth from the funiculus, often pulpy; hence arillar, arillate.

aristate: with a stiff awn or seta.

articulate: jointed; with nodes or joints where parts may move or separate.

artificial: modified by humans.

ascending: directed upwards, usually at a sharp angle.

attenuate: gradually tapering.

auricle: an ear-shaped appendage or lobe; hence auriculate.

autogamous: automatic self-pollination within a fl.

autotrophic: see holophytic.

awn: a stiff or bristle-like projection often from the tip or back of an organ.

axil: the upper angle between 2 dissimilar parts; hence axillary, occuring in the axil.

axis: the main stem or central longitudinal support; plural axes.

baccate: of fr., berry-like, with pulpy mesocarp.

barbs: retrorse projections, particularly on hairs; hence barbate; diminutive barbellate.

basifixed: attached by the base; usually of anthers.

basiscopic: of a pinna or lobe pointing toward the base.

beak: a prominent extension of an organ, e.g. of an achene.

berry: a fleshy fr. containing several to many seeds but not a bony endocarp.

biennial: living usually 2 years and flowering and dying in the second.

bifarious: arranged in 2 opposite rows.

bilabiate: with 2 lips.

blade: the expanded part of a lf, petal, etc.

bole: the unbranched trunk of a tree.

bract: a modified, often much reduced lf, especially the small scale-like lvs of an infl.; diminutive bracteole; hence bracteate, bracteolate.

bristle: a stout, stiff hair.

bud: a much condensed, undeveloped shoot or fl.

budding: grafting by inserting a bud under the bark of the stock; hence budded.

bulb: a thickened, compressed stem with ± fleshy, scale-like lvs; diminutive bulbil; hence bulbous.

bullate: with the surface blistered or puckered.

bush: a shrub with dense branches, resulting usually in dense habit.

caducous: falling off at an early stage.

caespitose: growing in tufts.

calli: callus deposits, usually appearing as small coloured dots, as on lvs of Oxalis.

callus: a hardened, usually thicker part; hence callosity.

calyx: the outer, usually greenish, whorl of floral parts, consisting of free or united sepals; plural calyces.

calyptrum: a hood or lid usually formed by the calyx and usually falling as a unit; hence calyptrate.

cambium: a generative layer of thin cells whose divisions produce rows of vascular tissue cells or cork cells.

campylotropous: of ovules, curved with the micropyle nearly meeting the funiculus.

canescent: with grey-white pubescence.

capillary: hair-like.

capitulum: a dense head-like infl. of usually numerous sessile fls, as in most Asteraceae; plural capitula; hence capitate.

capsule: a dry dehiscent fr. formed by the union of 2 or more carpels.

carnivorous: of plants that capture and absorb animals.

carpel: one unit of the ♀ part of the fl., consisting of an ovary, style, and stigma; hence carpellate.

carpophore: a prolongation of the floral axis above the level of attachment of the perianth, either supporting the gynoecium or, as in most Apiaceae, carrying the pendulous ripe mericarps.

cartilaginous: firm and tough but flexible, like cartilage.

caruncle: a callus-like appendage at or about the hilum of a seed, often paler in colour than seed coat; hence carunculate.

casual: occuring sporadically and not establishing.

catkin: a spike or spikelike infl., usually unisexual, on a pendulous to erect axis.

caudate: bearing a tail-like appendage.

caudex: a stem with subsidiary organs, usually of herbaceous plants.

caulescent: having an evident above-ground stem.

cauliflorus: producing fls from old wood, especially the trunk.

cauline: belonging to the stem, especially its upper aerial part.

caulorrhiza: woody rootstock, as in Geraniaceae.

chartaceous: of thin papery texture, not usually green.

chasmogamous: of fls, opening for pollination.

choripetalous: with separate petals.

cilia: short hairs forming a fringe, usually at a margin; hence ciliate; diminutive ciliolate.

circinate: coiled, the apex innermost, as in young fern fronds.

circumscissile: dehiscing by a circular horizontal suture, the top valve coming off as a lid.

clathrate: latticed or pierced by apertures.

clavate: club-shaped, thickened toward the apex.

claw: a narrow stalk-like base of a petal or sepal.

cleistogamous: of fls, remaining closed and self-fertilizing; hence cleistogamy.

clone: a vegetative propagule, genetically identical with the parent; hence clonal.

coalescent: of the incomplete fusion of like parts, as in many compound frs.

cocci: the parts of a dry lobed fr., each part 1-seeded.

coherent: like parts in close contact but not fused.

column: body formed by the union of stamens, style and stigma, or sometimes of staminal filaments only; especially in Stylidiaceae.

coma: a tuft of hairs, as in seeds of Epilobium.

commissure: the face by which one carpel joins another.

compound: composed of several ± similar parts, as opposed to simple; as in compound lvs.

concolorous: of ± the same colour throughout.

conduplicate: folded together lengthwise, along the midrib, with the upper surface within; of cotyledons folded lengthwise around an incumbent radicle.

cone: a general term for the hardened fructifications, especially of the Pinaceae and Cupressaceae; hence coning.

conelet: a cone up to the time of fertilization.

confluent: blending or running together.

connate: joined together, especially of like parts.

connective: the tissue connecting the 2 lobes of an anther.

connivent: converging together; usually of 2 or more organs with their bases separated and their apices approaching one another.

contiguous: touching, adjoining.

contractile: capable of expanding and contracting, usually of roots.

convolute: rolled together, longitudinally; usually of floral whorls in bud, each part overlapping next unit of whorl.

cordate: heart-shaped, with the notch at the base.

coriaceous: of somewhat leathery texture, tough.

corm: a short, swollen, underground stem, usually erect and invested by few scales and lasting only one season.

corolla: the inner, often showy, whorl of floral parts, consisting of free or united petals.

corona: appendage projecting upwards or inwards from corolla, usually crown-like; hence coronal.

corrugate: evenly furrowed or wrinkled.

corymb: an indeterminate flat or convex-topped infl.; hence corymbiform, corymbose.

cotyledon: seed lf, usually present on embryo in the seed.

crenate: with shallow rounded teeth, the sinus acute; diminutive crenulate; hence crenation.

crepuscular: occurring in late afternoon and early evening, especially of fl. opening.

crispate: with margins curled, densely undulate.

croziers: young uncurling fern fronds.

cucullate: hooded or hood-shaped

cultigen: a plant known only in cultivation.

cultivar: a selected and named form reproduced in cultivation but not necessarily referable to a botanical sp.

cultrate: shaped like a knife blade.

cuneate, cuneiform: wedge-shaped, gradually and evenly narrowed to base.

cup: a cup-like structure; diminutive cupule; hence cupuliform, cupular.

cusp: a sharp rigid point; hence cuspidate.

cuticle: the outermost, ± impermeable layer on the epidermal cells.

cyathium: a cup-like structure, especially the perianth-like involucre surrounding fls in Euphorbia.

cyme: a determinate, usually flat or convex-topped infl.; hence cymose.

cypsela: an achene bearing an adnate calyx, as in some Asteraceae and related families.

deciduous: falling at end of season or when function has ceased; often of lvs in autumn.

declinate: bent or curved downwards.

decumbent: lying along the ground with the tip ascending.

decurrent: running or extending downwards, especially of lvs where the base is prolonged down the axis and adnate to it.

decussate: of lvs or other organs that are opposite with successive pairs at right angles and so forming 4 rows, as in lvs of some Crassula spp.

dehiscent: opening, usually regularly, to shed contents when ripe; hence dehiscence.

deltate: broadly triangular.

dendroid: resembling a tree in shape or mode of branching.

dentate: with sharp teeth perpendicular to margin, the sinus ± open; diminutive denticulate.

denticle: a minute tooth.

depressed: flattened from above.

diadelphous: in 2 sets, as of stamens in most papilionoid Fabaceae where 9 are united and one is free.

dichasium: a cyme in which the branches are opposite and ± equal in length, see Fig. 51.

dichotomous: forking into 2 equal branches.

didymous: twinned, the 2 parts similar and attached by a short length of their inner surface.

didynamous: with 4 stamens, one pair long, one pair short.

digitate: spreading from the centre like the fingers of a hand.

dioecious: the population comprising 2 sexual morphs, these being almost or quite strictly ♂ and ♀; hence dioecism.

diploid: having 2 sets of chromosomes.

disc: a flattened, ± fleshy development of the receptacle or coalescent nectaries, or the non-ligulate part of the capitulum made up of tubular fls in Asteraceae; hence discoid.

discolorous: of 2 different colours, especially of upper and lower lf surfaces.

discrete: separate, not adherent or coherent.

distal: towards the free, as opposed to the attached or proximal, end of an organ.

distichous: arranged in 2 opposite rows so as to lie in one plane.

distylous: having a floral dimorphism in which the anthers and stigmas vary in height between morphs so that the sequence of heights is different in each morph.

diurnal: occurring during the day, especially of fl. opening.

divaricate: spreading at a very wide angle.

domatia: small pits on the adaxial surface of lvs, between the veins and midrib or near the margin.

dorsally: relating to the back.

dorsifixed: attached by the back, especially of anthers.

drupe: a "stone fr.", the seed enclosed in a bony endocarp which is surrounded by a ± fleshy mesocarp; hence drupaceous; diminutive drupelet.

e- or ex-: prefix meaning lacking, e.g., ebracteate, eglandular, exstipulate.

elater: slender thread-like or coiled body found in sporangia and connected with spore dispersal.

ellipsoid: of a solid object, elliptic in section or outline.

elliptic: in the shape of an ellipse, rounded at both ends, widest in the middle.

emarginate: with a shallow notch at the apex.

embryo: the rudimentary plant within a seed, usually developing from a zygote.

endemic: native only to a particular country or region, in this Flora to N.Z. unless otherwise stated.

endosperm: the nutritive tissue outside the embryo in a seed.

endosporic: with the gametophyte wholly or partly growing within the spore wall.

endothecial: of the inner lining of the loculus of an anther.

entire: with a continuous margin completely lacking teeth.

entomophilous: pollinated via insects.

ephemeral: very short-lived.

epicalyx: a calyx-like structure immediately below the true calyx.

epicarp: of frs, the outer layer of the pericarp.

epicormic: of buds and shoots developing from the trunk.

epidermis: the outermost cell layer of most organs.

epigeal: above ground, especially of cotyledons.

epigynous: borne on the ovary; of a fl. in which stamens and perianth arise level with or above the top of the ovary which is adnate to the surrounding receptacle.

epimatium: in Podocarpaceae, a scale ± fused with the integument and partly or completely surrounding the ovule.

epipetalous: borne on the petals or corolla.

epiphyte: a plant growing upon another but not organically connected with it; hence epiphytic.

episepalous: borne on the sepals or calyx.

ericoid: of habit or lvs, with small, crowded, ± narrow-linear lvs, as in many Erica spp.

erose: with an irregular margin as if gnawed.

eusporangiate: with each sporangium originating from a group of cells, its wall > 1 cell thick.

evergreen: having lvs all through the year.

exfoliating: to peel off in strips or plates, as with bark from a trunk.

exocarp: the outer layer of the pericarp or fr. wall.

exosporic: with the gametophyte free-living, not enclosed by the spore wall.

extrorse: directed outward, used of anther dehiscence.

facultative: optional.

falcate: sickle-shaped; strongly curved.

farina: a mealy covering; hence farinose.

fasciation: an abnormal widening and flattening of the stem.

fascicle: a close cluster or bundle; hence fascicled, fasciculate, fascicule.

fastigiate: with the branches ± erect and close to the axis.

female: plants, fls, or fl. parts bearing the megagametophytes or ovules.

fenestrated: with window-like openings or translucent areas.

ferrugineous, ferruginous: rust-coloured.

-fid: suffix denoting cleft, e.g., 3-fid, cut into 3 narrow lobes.

filament: thread-like organ, especially the stalk bearing the anther; hence filamentous.

filiform: thread-like.

fimbria: hair or process forming part of a fringe; plural fimbriae; hence fimbriate.

fistular, fistulose: hollow and cylindric.

flabellate: fan-shaped.

flaccid: limp, not rigid.

flexuous, flexuose: having a wavy or zigzag form.

floccose: clad in tufts of soft woolly hairs, usually easily removed.

floral: of or pertaining to the fl.

floret: a small fl., used especially of individual fls in a composite head.

floriferous: flower-bearing.

flower: a complex of organs performing the reproductive functions of a flowering plant, usually comprising one or more stamens and pistils, surrounded by a corolla and a calyx.

foetid: having an offensive or strong odour.

foliaceous: leaflike.

foliose: bearing numerous or crowded lvs.

follicle: a dry dehiscent fr. formed from one carpel and dehiscing along ventral side; hence follicular, folliculate.

forma: a minor variant of a sp., perhaps differing by only a single character or caused by a single gene, and often occurring sporadically along with other forms of the sp.; plural formae.

free-central: of a placenta arising from the base of a 1-locular ovary but not touching the apex or the sides.

frond: a lf, used especially of ferns.

fruit: the ripened ovary containing the seeds; often used to include associated fleshy parts of the receptacle.

fruticose: shrubby.

fugaceous: falling very early, as of some petals and sepals.

fulvous: tawny; dull yellow with a mixture of grey or brown.

funicle, funiculus: the stalk of an ovule attaching it to the ovary wall or placenta.

funnelform: with a tube gradually widening upwards.

fused: joined together.

fusiform: spindle-shaped; of a solid ± swollen in the middle and narrowed to both ends.

gall: a swelling produced usually by an insect parasite.

gametophyte: the haploid body produced by the germinating spore and bearing sexual organs; in lycopods and ferns a small separate plant.

gamopetalous: with the petals united, at least at the base.

gamophyllous: with lvs or floral units united by their edges.

gamosepalous: with the sepals united, at least at the base.

gelatinous: jelly-like.

gemmae: asexual propagules, appearing like buds.

geniculate: with a knee-like bend.

genotype: genetic constitution, or a particular form so determined.

gibbous: somewhat swollen on one side, usually near the base, forming a pouch or sac.

glabrate: almost glabrous.

glabrescent: becoming glabrous.

glabrous: without hairs of any sort.

gland: a secreting organ or part, the secretion oil, resin, or other liquid; hence glandular.

glaucous: a distinct bluish green colour, not necessarily caused by bloom; hence glaucescent, slightly glaucous or becoming so.

globose, globular: of a solid object, spherical.

glochid: a small, barbed spine, as in Acaena; plural glochidia; hence glochidiate.

glomerule: a very dense cluster.

glutinous: sticky, or covered with sticky exudate.

granular, granulate, granulose: covered with small grain-like bodies.

gynobasic: of a style appearing to be inserted at the base of an ovary.

gynodioecism: the population comprising 2 sexual morphs, one being strictly ♀, the other, termed ♂, producing some seed; hence gynodioecious.

gynoecium: the ♀ part of the fl. made up of one or more carpels.

gynomonoecious: having ♀ and ⚥ fls on the same plant.

gynophore: a stalk raising the base of the ovary above the level of attachment of the androecium.

gynostegium: a column formed by the fusion of the stamens and the style and stigma; hence gynostegial.

halophyte: a plant growing within the influence of salt water.

hardy: able to withstand cold temperatures, especially of frost-tolerance.

hastate: shaped like an arrowhead, with the basal lobes pointed or narrow and spreading at a wide angle.

haustoria: the absorbing organs of a parasite or hemiparasite.

head: a short, dense infl. of sessile or almost sessile fls.

helicoid: of cymes modified by the loss of fls from the same side of each dichotomy, hence curving, common in Boraginaceae.

hemispheric: of solid objects, like 1/2 a globe.

herb: any vascular plant that is not woody; hence herbaceous.

herbal: a book describing plants used for food, flavouring, scent or medicine; hence herbalist, one with knowledge of such plants.

herbarium: a collection of dried or preserved plants, systematically arranged; plural herbaria.

herbfield: vegetation type, usually closed and distinguished by the abundance of herbs.

hermaphrodite: having both stamens and carpels.

heterogamous: with 2 or more kinds of fls, e.g., of the heads of many Asteraceae where ray florets differ from disc florets.

heteromorphic: with 2 or more forms.

heterophyllous: bearing lvs of 2 or more forms, especially of plants with aquatic and aerial lvs.

heterosporous, heterosporus: producing 2 or more kinds or sizes of spores.

heterostylous: having a floral polymorphism in which the anthers and stigmas vary in height among morphs so that the sequence of heights is different in each morph.

heterotrophic: of plants which are entirely saprophytic.

hilum: the scar on the seed marking the place of attachment to the funiculus.

hispid: bearing stiff, ± bristle-like hairs; diminutive hispidulous.

holophytic: of plants maintained by own organs, not dependent on other plants - free-living.

homogamous: with one kind of fl. only, especially used of some Asteraceae.

homologous: of the same origin, but often differing in form or function.

homomorphic: of one form.

homonym: a name spelt exactly the same as another but applying to a different type specimen.

homosporous: producing spores of one kind or size only.

horizontal: positioned at right angles to the main axis.

host: a plant having a parasite living upon it.

husk: an outer, usually loose, covering of some frs.

hybrid: a plant resulting from a cross between parents which differ sufficiently to be accorded taxonomic recognition, the hybrid name usually preceded by, or names of the parents joined by, ×; hence hybridise, hybridism.

hydrocyanic: producing hydrogen cyanide, as in lvs of some Fabaceae.

hydrophytic: living in water.

hygrochastic: of a capsule opening when wet and closing when dry, as in Aizoaceae.

hypanthium: a cup-like structure produced by the fusion of the bases of the floral envelopes and androecium, often referred to as a calyx tube.

hypodermal: the layer of cells immediately under the epidermis.

hypogeal: especially of cotyledons that do not appear above ground.

hypogynous: borne below the ovary; of a fl. in which perianth and stamens arise below the ovary.

imbricate: overlapping, like roof-tiles; in buds, with the edges of the organs overlapping, but not regularly.

immersed:± imbedded.

imparipinnate: pinnate with a single terminal pinna or leaflet.

inaperturate: without apertures, usually of pollen grains.

incision: a deep sharp cut; hence incised.

included: not projecting beyond the enveloping structure, as of stamens included within the corolla tube.

indehiscent: not opening to release its contents, except finally by decay.

indument, indumentum: general terms for a covering of hairs of any form.

induplicate: with the edges folded or turned in.

indurate: hardened and toughened.

indusium: an outgrowth of tissue ± covering the sorus in some ferns; plural indusia; hence indusial.

inferior: situated below another organ or part; an inferior ovary is surrounded by and fused with the receptacle and so is below the insertion of the perianth.

inflorescence: a general term for the branch system on the floral axis, or for the arrangement of fls within this.

infructescence: a general term for the fruiting structure formed from an infl.

inserted: attached to or growing upon.

integument: a covering; used especially of the covering of the nucellus of an ovule.

internode: the part of an axis between 2 nodes.

interpetiolar: between the petioles; often applied to coalescent stipules of opposite lvs.

interstitial: placed in the space between 2 other organs.

interveinal: between the veins.

intramarginal: position just within the margin.

intrapetiolar: position on the inside of the petiole.

intrastaminal: positioned within the stamens, often of a floral disc.

introgression: infiltration of the genes of one taxon into the genotype of another through crossing and backcrossing.

introrse: facing inwards or towards the axis; often applied to anthers that open towards the centre of a fl.

invagination: the pushing inwards of a part of layer of cells so as to form a pocket.

investment: an enclosing covering or envelope; hence invested.

involucre: one or more whorls of bracts or lvs, often calyx-like, surrounding an aggregation of fls; hence involucral, involucrate; diminutive involucel.

involute: rolled inwards or to the upper (adaxial) side.

irritable: highly responsive to touch.

isobilaterally: with the 2 sides similar.

isodiametric: having equal diameter, particularly of structures of polyhedral form.

isozyme: the product of a particular allele at a heterozygous locus.

keel: a sharp central ridge, like the keel of a boat; used also of the united anterior petals of papilionoid Fabaceae.

kernel: the softer, often edible, part within the shell or harder outer layer of a fr.

lacerate: irregularly torn or cleft.

laciniate: deeply, usually irregularly, divided into very narrow, pointed segments.

lamellate: of placentation where a compound ovary comprises many carpels and the partitions have the ovules scattered over their surfaces.

lamina: a thin, flat organ or part, especially the expanded portion or blade of a lf; plural laminae, laminas.

lanate: clad in woolly, usually intertwined, hairs.

lanceolate: lance-shaped, much longer than wide and more gradually tapered to apex than to base.

lateral: positioned on the side or in an axil, as opposed to terminal.

latex: milky juice.

latiseptate: of a flattened 2-locular capsule (or silique) with septum across the broad diameter.

layer: stems covered with earth and then rooting; hence layering.

leader: a vigorous dominant shoot.

leaf: an expanded organ produced laterally from the stem and usually comprising the blade (lamina), petiole and stipules.

leaflet: one element of a compound lf.

legume: a simple, 1-celled and usually 2-valved fr.; also used as a name for members of the Fabaceae.

lenticel: a corky spot on young bark functioning as a pore; hence lenticellate.

lenticular: the shape of a ± circular, biconvex lens.

lepidote: clad in scurfy scales.

leptosporangiate: with each sporangium originating from a single superficial cell, its walls consisting of a single layer of cells.

liane: a woody climbing plant; hence lianoid.

ligule: a strap-shaped body, as the limb of the corolla in many Asteraceae; hence ligular, ligulate.

limb: the expanded part of a petal or sepal.

linear: very narrow and with parallel margins.

lingulate: tongue-shaped.

lobe: a recognisable but not separate division of a lf, calyx or corolla, especially when rounded; hence lobate; diminutive lobule, lobulate.

locule, loculus: a compartment or cavity of an organ, especially an ovary, fr. or anther; plural locules, loculi; hence locular.

loculicidal: of dehiscence by the splitting of the outer wall of each loculus.

loess: a fine grain deposit of yellowish grey loam, usually wind blown.

lomentaceous: of a legume or pod strongly constricted between the seeds.

lunate: with the shape of a crescent moon.

lurid: dingy brown or dull yellow.

lustrous: glossy.

lyrate: obovate with several deep recesses or sinuses on each side which gradually diminish in size to the base.

Macaronesia: the islands off the coast of N. Africa - Madeira Is, Canary Is, and Azores.

macrosporangium: a sporangium containing macrospores, the larger kind of spores in heterosporous plants.

male: plants, fls, or fl. parts bearing the microgametophytes or pollen.

mammillate: with nipple- or teat-shaped projections.

mangrove: subtropical or tropical intertidal forest or scrub.

mealy: covered with or as if with a fine dust or powder.

medifixed: attached at the centre, especially of anthers and hairs.

megaphylls: lvs with a branching vein system.

megasporangium: a sporangium containing macrospores, the larger kind of spores in heterosporous plants.

megaspore: the larger kind of spore in heterosporous plants.

megasporophyll: a ± modified leaflike structure bearing megasporangia; in angiosperms the carpel.

membranous: thin and ± pliable.

mericarp: a dry 1-seeded part of a fr., usually 1/2 of a schizocarp as in Apiaceae.

-merous: denoting parts and used mainly of floral organs, e.g., 5-merous, having the sepals, petals, etc. in whorls of 5.

microphylls: lvs with an unbranched vascular strand.

microsporangium: a sporangium containing microspores, the smaller kind of spores in heterosporous plants.

microspore: the smaller kind of spore in heterosporous plants.

microsporophyll: a ± modified leaflike structure bearing microsporangia; in angiosperms the stamen.

microsp.: a sp. in a genus where numerous spp. are recognised by minor characters, the entities usually maintained only via apomixis or selfing, as in Hieracium, Rubus fruticosus.

monadelphous: in one group, as of stamens all united by their filaments into a tube or column.

moniliform: constricted at short intervals and so appearing like a string of beads.

monocarpic: flowering and fruiting only once, usually after several years, and then dying.

monochasium: a 1-branched cyme, usually resulting from reduction of laterals, sometimes appearing raceme-like, see Fig. 51; hence monochasial.

monoecious: with unisexual fls, ♂ and ♀ on the same plant.

monolete: having a single scar or fissure on the spore wall, characteristic of bilateral pteridophyte spores.

monostelic: having one stele or central cylinder of vascular tissue.

monotypic: having only one representative, as of a genus with one sp.

monstrosity: horticultural novelties resulting from abnormality of growth, e.g., double fls.

morph: a discrete form, often with a genetical basis.

mouth: the opening at the apex of a tube, particularly of corolla or calyx.

mucilage: a viscous fluid; hence mucilaginous.

mucro: a short sharp tip or extended midrib; hence mucronate; diminutive mucronulate.

multicipital: with many heads, referring especially to the stock of a single root bearing many short branches.

muricate: rough owing to many minute, hard outgrowths; diminutive muriculate.

naked: of a part lacking bracts or appendages or of a fl. lacking a perianth.

napiform: turnip-shaped.

nectary: gland exuding fluid containing sugars and other compounds as a reward for pollinators; hence nectar, nectariferous.

neuter: of a fl. lacking functional sexual organs.

nocturnal: occurring during the night, especially of fl. opening.

node: a place on a stem marked by one or more lvs; hence nodal.

nut: an indehiscent, 1-seeded fr. with a hard, woody wall; diminutive nutlet, sometimes applied to the hard seed-like divisions of a fr. as in Boraginaceae.

nyctinastic: nocturnal folding of lvs or movement of other parts.

ob-: signifies inversion, e.g., obovate, obcordate.

obconic: like an inverted cone.

obligate: of a plant not able to adopt a different habit, e.g., obligate apomict, obligate parasite.

oblique: with sides unequal or meeting unequally.

obpyramidal: like an inverted pyramid.

obtriangular: like an inverted triangle.

obtuse: blunt.

ochrea: a tubular sheath at the node formed by the fusion of 2 (usually membranous) stipules, as in Polygonaceae; plural ochreae.

offset: a new plant produced vegetatively from a lateral shoot.

opaque: applied to a surface, dull not shining.

opercula, operculum: a lid or cover separated by a ± circular line of division, as in the fr. of Plantago.

opposed: placed opposite another organ.

opposite: of a pair of organs arising at the same level on opposite sides of a stem, especially of lvs.

orbicular: circular or nearly so.

orthotropous: of ovules, with axis in a straight line with the funiculus, and so erect, the micropyle remote from the placenta.

ostiole: a small opening or mouth, as in the infl. of Ficus.

oval: broadly elliptic, narrowing from the middle to rounded ends.

ovary: the part of the gynoecium that encloses the ovules.

ovate: egg-shaped, attached by the broad end.

ovoid: of a solid body with an ovate outline.

ovule: the structure which contains the megagamete or egg and after fertilization develops into the seed; ovuliferous, bearing ovules.

palate: a prominent part enclosing the throat of a corolla, especially in Scrophulariaceae.

palmate: radially divided into separate parts and so compound.

palmatifid, palmatisect: radially divided or dissected but not compound.

pandurate: fiddle-shaped, ± obovate but with a waist.

panicle: an indeterminate, branched infl. with pedicellate fls; hence paniculate.

papilionaceous: butterfly-like; of a corolla with the parts in the form and arrangement of those of a sweet pea.

papilla: minute, pimple-like process; hence papillate, papillose.

pappus: a persistent calyx of special form crowning the achene of most Asteraceae; parts are usually numerous, simple or feathery hairs or bristles, or sometimes scales.

parallelogrammoid: with 4 sides, the opposite sides parallel.

paraphyses: sterile filaments or hair-like structures of various shapes occurring amongst sporangia in a sorus.

parasite: a plant attached to and deriving nourishment from another living plant; hence parasitic.

parietal: pertaining to the wall; placentation is parietal when the ovules are borne on the walls of the ovary or on minute projections therefrom.

paripinnate: evenly pinnate, i.e., without a terminal leaflet.

partite: deeply divided.

pectinate: divided in a comb-like fashion, with the segments narrow and close-set.

pedate: foot-like, with the segments arising from a broader base than palmate.

pedicel: the stalk of an individual fl.; hence pedicellate.

pendulous: drooping.

peduncle: a common leafless axis bearing several fls; hence peduncled, pedunculate.

pellucid: clear, transparent or nearly so.

peltate: shield-like, with the stalk attached well inside the margin.

pelviform: basin-shaped.

penicillate: provided with a brush of hairs.

penninerved: pinnately veined.

perennial: with a life span of > 2 years; hence perennate.

perfoliate: of a sessile lf or bract completely encircling the axis, so having the stem apparently passing through it.

perianth: the floral envelopes considered as a whole; especially used when the corolla and calyx are not well differentiated, or when either one is absent.

pericarp: the wall of the ripened ovary, i.e., of the fr.

perigynous: surrounding the ovary; of a fl. in which perianth and stamens arise from the edge of a ± cup-shaped receptacle, surrounding but free from the ovary.

perisperm: food reserve in a seed, formed from the nucellus.

perulae: scales, usually those covering a seed or bud.

petal: a unit of the corolla, when completely free, usually coloured other than green; hence petaline; petaloid, resembling a petal.

petiole: the stalk of a lf; hence petiolar, petiolate.

petiolule: the stalk of a leaflet; hence petiolulate.

phenotypes: the sum of the characteristics manifested by an organism - varying as a result of the genotype or environment.

phyllode: a widened petiole or rachis functioning as a lf blade; hence phyllodinous.

phytophagous: plant-eating.

pilose: bearing soft shaggy hairs.

pinnate: compound with the parts arranged on either side of an axis; pinna, pinnule, the elements of a pinnate lf.

pinnatifid, pinnatisect: divided or dissected to varying degrees in a pinnate fashion but not compound.

pistil: the gynoecium, comprising the ovary, style and stigma.

placenta: the place or part of an ovary where the ovules are attached.

placentation: the arrangement of the placentae within the ovary, i.e., axile, free-central, parietal.

plano-: having one surface flat, as in plano-convex.

plicate: folded into pleats, usually lengthwise.

ploidy: of the level of chromosome duplication.

plumose: feather-like.

pneumatophores: specialised root branches produced by some plants growing in water, especially mangroves, the branches erect and projecting into the air.

pollination: the process by which pollen is transferred from anthers to stigmas.

pollinator: an agent used by a plant to achieve pollination and fertilization.

pollinium: a coherent mass of pollen, as in Asclepias; plural pollinia.

polymorphic: having several forms, usually of spp. varying greatly among individuals; hence polymorphism.

polyphyletic: of a group of taxa classified together even though some have quite distinct evolutionary histories.

polyploid: having a chromosome complement of 2 or more times the normal diploid number.

polystelic: having more than one stele or central cylinder of vascular tissue.

pome: a simple, fleshy fr. with a soft exterior and papery or cartilaginous carpels, e.g., an apple.

poricidal: opening or dehiscing by pores.

posterior: on the side nearest the axis and so appearing at the back.

pouch: a bag-like cavity.

prickle: a small sharp outgrowth of superficial origin; diminutive pricklet.

primocane: the vigorous, first year non-flowering stems of Rubus.

procumbent: lying ± flat along the ground but not rooting.

prophyll: a small lf or bract.

prostrate: a general term for lying flat on the ground.

protandrous: with the anthers shedding pollen before the stigma is receptive.

prothallus: a small body produced by the germinating spore and bearing sexual organs, the gametophytic generation of pteridophytes.

protogynous: with the stigma receptive before pollen is shed from the anthers.

provenance: derivation as to region.

proximal: towards the attached, as opposed to the free or distal, end of an organ.

pruinose: bearing a waxy or powdery bloom on the surface.

pseudo-: false.

puberulent, puberulous: covered with exceedingly fine, short, dense hairs.

pubescent: clad in short soft hairs; hence pubescence.

pulvinate: of cushion-like form; growing in dense cushions.

punctate: with dot-like markings, frequently owing to translucent or coloured glands; hence punctiform; diminutive puncticulate, punctulate.

pyrene: a seed with a bony endocarp, usually when there are several in a fleshy fr.

pyriform: pear-shaped.

quadrate: square or rectangular.

raceme: an unbranched, ± elongate, indeterminate infl. with stalked fls; hence racemoid, racemose.

rachis: the axis of an infl. or of a compound lf.

radiate: diverging from or arranged around a common centre, as the spokes of a wheel; also bearing ray florets, as in many Asteraceae.

radical: of lvs, arising from the stock or crown of a root or from a rhizome.

radicle: the primary root of an embryo.

ramiflorous: producing fls from branches.

raphides, rhaphides: bundles of crystals of calcium oxalate.

ray: the florets with strap-like corollas in many Asteraceae; also a peduncle forming part of an umbel.

receptacle: the ± expanded termination of the stalk which bears the fl. parts, the fl. or the fl. head.

recurrent: of flowering, continuing to bloom through the season after the initial flowering period, as in many roses.

reflexed: bent sharply backwards.

regma: a dry dehiscent fr. of usually 3 (rarely more) cells as in Euphorbiaceae.

regular: symmetric; of fls radially symmetric = actinomorphic.

reniform: kidney-shaped.

replum: a frame-like placenta from which the valves fall away in dehiscence, as in many Brassicaceae.

resinous: containing or producing resin, and so sticky.

resupinate: twisted through 180°.

reticulate: in the form of a network; hence reticulum.

retrorse: bent backwards or downwards, usually abruptly.

retuse: with the apex rounded with a small notch.

reversion: of shoots bearing lvs of juvenile form.

revolute: rolled outwards or to the lower (abaxial) side.

rhizome: an underground stem, usually spreading ± horizontally; hence rhizomatous.

rhombate, rhombic: diamond-shaped; hence rhomboid, similar but in 3 dimensions.

root: the descending axis of the plant, initially developing from the radicle.

rootcrown: the part of the stem at the ground surface.

rootstock: a short, erect underground stem; a stem with roots, used for grafting.

rosette: a group of organs radiating from a centre, especially used where internodes are short and lvs overlap; hence rosulate.

rostrate: beaked.

rotate: wheel-shaped; of a gamopetalous corolla with a flat, circular limb at right angles to a short tube.

rotund: orbicular.

rudimentary: imperfectly developed, vestigial.

rufous: of reddish colour.

rugose: wrinkled; hence rugosity; diminutive rugulose.

ruminate: of irregular mottled appearance.

runcinate: pinnately and rather sharply lobed, with the lobes ± directed backwards.

runner: a slender, ± prostrate or arching lateral stem rooting at the nodes.

rupestral: growing on rocks.

sac: a pouch-like structure; hence saccate.

sagittate: in the form of an arrowhead with the basal lobes at a narrow angle to the stalk.

saline: salty.

salverform: of a gamopetalous corolla with a ± long slender tube abruptly expanded into a flat limb.

samara:1-seeded winged fr.

saprophyte: a plant that obtains its food from dead organic matter; hence saprophytic.

scabrid, scabrous: rough to the touch because of minute harsh projections; diminutive scaberulous, scabridulous.

scale: a small ± leaflike organ, often dry and membranous.

scandent: climbing, usually with special climbing organs.

scape: a leafless, ± elongate peduncle arising from the crown, with or without scales or bracts, usually with a single fl. by reduction; hence scapose.

scarious: very thin, dry, and ± translucent.

schizocarp: a dry fr. splitting into 1-seeded halves when ripe.

sclerotic: hardened, stony in texture.

scorpioid: of a cymose infl., with the young axis coiled like the tail of a scorpion.

scrambler: an unspecialised climber, not usually reaching a great height.

scree: loose, ± moving, stony debris.

scutate: shield-shaped.

secund:1-sided, with all the fls, lvs or other parts appearing to be arranged along one side of the axis.

seed: the reproductive body formed usually from the fertilized ovule.

segment: an individual free part of an organ.

selfing: of pollination and fertilization within one plant.

sensitive: responding to stimulation, particularly touch as in triggered stamens of Berberis.

sepal: one separate part of a calyx of free members, especially when green and leaflike.

septicidal: dividing along a septum; of dehiscence by breaking apart of the fused carpel walls that form the septa between loculi.

septifragal: when in dehiscence the valves break away from the septa.

septum: a partition or cross-wall; plural septa, septae; hence septate.

seriate: arranged in series or rows, e.g., 1-seriate, in 1 row.

sericeous: silky; clad in soft, simple, ± appressed hairs.

serrate: sharply toothed with the teeth pointing forward; hence serration; diminutive serrulate.

sessile: without a stalk.

seta: a fine bristle-like structure; hence setaceous, setose; diminutive setulose.

shrub: a woody plant with many stems and lacking a distinct trunk; diminutive shrublet, a very small shrub, usually with slender stems.

siliceous: containing minute particles of silica.

silique: a capsule, usually 2-locular, with 2 valves falling away from a frame (replum) bearing the seeds, characteristic of Brassicaceae; hence siliquiform; diminutive silicle, used when length is not or little > 3× width.

simple: not divided into several ± similar separate parts; contrasted with compound.

sinuate: with shallow broad waves to the margin but not undulate.

sinus: the recess between 2 lobes or segments.

solitary: occurring singly.

sordid: of an impure or dirty white appearance, especially of the pappus of some Asteraceae.

sorus: a cluster of sporangia, as in most ferns; plural sori.

spathe: a large bract ± enclosing an infl.

spathulate: spoon-shaped.

spherical: of a solid body with a circular outline.

spike: an unbranched, ± elongated, indeterminate infl. with sessile fls; hence spicate, spicately, spiciform; diminutive spikelet.

spine: a stout, woody process with a sharp point; hence spinescent, spinose, spinous; diminutive spinules, spinulose.

sporangiophore: a stalk bearing a sporangium.

sporangium: a sac or other structure containing spores; plural sporangia.

spore: a simple, asexual, usually 1-celled reproductive body, as in ferns.

sporocarp: a thick-walled body containing sporangia.

sporophyll: a ± modified leaflike structure bearing sporangia.

sporophyte: the plant which bears not sexual organs but asexual spores; in lycopods and ferns the ordinary plant; in seed plants the whole plant except the embryo sac and pollen grain.

sport: a plant or part of a plant exhibiting abnormal variation, often as a result of somatic mutation, often used to produce new cvs.

spreading: extended laterally, of trees with broad crown.

spur: a sac or tubular part, often nectar-secreting, of a petal or sepal; hence spurred.

stamen: the pollen-bearing organ, composed of an anther with pollen sacs and its supporting filament.

staminode: a non-functional stamen, sometimes petaloid.

stellate: star-shaped, usually of hairs with many radiating arms.

sterile: not producing seed, spores or pollen capable of germination.

stigma: the part of the carpel that is receptive to pollen, usually found at or near the tip of the style; hence stigmatic.

stipe: a stalk; mainly used of petiolar part of fern lf; hence stipitate.

stipel: the stipule subtending a leaflet; hence stipellate.

stipule: one of a pair of scale-like or leaflike appendages at the base of a petiole; hence stipular, stipulate.

stolon: a stem, ± horizontal or arched or running along the ground, rooting and capable of forming a new plant at its tip; hence stoloniferous.

stomata: pores in the epidermis through which gases diffuse.

striate: with fine longitudinal lines or ridges.

strigose: with sharp, ± appressed hairs, often ± swollen at base; diminutive strigulose, strigillose.

strobilus: a cone-like structure containing reproductive organs, as in lycopods and pines; plural strobili; hence strobilate.

strophiole: an aril-like but hard appendage of a seed, at or near the micropyle; hence strophiolate.

style: the ± elongated part of the carpel between the ovary and stigma; hence stylar.

stylopodium: an enlarged basal part of a style, as in many Apiaceae.

sub-: prefix indicating somewhat less than.

subshrub: a small plant with stems ± woody towards the base.

subtend: stand below, but usually close to, another organ, as a bract to its fl.

subulate: awl-shaped; tapering from a wider base to a sharp apex, ± circular in cross-section.

succulent: fleshy, soft and thickened in texture.

sucker: leafy shoot of subterranean origin; hence suckering.

sulcate: with longitudinal grooves, less pronounced than furrowed.

superior: situated above another part; an ovary is superior when it is free from the receptacle, with the perianth and stamens inserted below or around it.

superposed: of a part or organ placed vertically above another, especially of buds.

suture: a line or seam formed at the junction of 2 margins; used especially of dehiscent frs.

sympatric: of taxa growing together in a particular region.

sympetalous: with the petals united at least at the base.

sympodial: of an axis made up of the basal portion of several branches, the apex of each branch dying or forming an infl. and growth continued by an axillary bud.

synangium: a concrescence of sporangia.

syncarp: an ovary of 2 or more united carpels; hence syncarpous, having united carpels.

synchronous: occurring at the same time.

syngenesious: with the anthers connate into a tube surrounding the style, as in the florets of Asteraceae.

synonym: a name based on the same type as an earlier published name, or based on a different type but the plant not considered distinct; hence synonymy, a list of synonyms.

talus: sloping mass of rock fragments, e.g., at foot of cliff.

taproot: the chief descending root.

tawny: brownish yellow, brownish orange, or tan-coloured.

taxon: a taxonomic group of any rank, e.g., family, genus, sp., var.; plural taxa.

tendril: a slender twining part of a climbing plant formed from the whole or part of a stem or lf; hence tendrilous.

tepal: an individual member of the perianth; hence tepaloid.

terete: circular in cross-section.

terminal: borne at the end of a stem.

ternate: compound with the parts arranged in threes; hence ternately.

ternatifid, ternatisect: divided or dissected in threes but not compound.

testa: the outer coat of a seed.

tetrad: a group of 4, in angiosperms particularly of pollen cells or pollen grains.

thalloid: of a plant body lacking differentiation into lf and stem.

throat: of gamopetalous corollas, the opening or mouth of the tube.

thyrse: a compact, ± cone-shaped panicle; hence thyrsoid.

tomentum: a dense, ± matted covering of soft hairs; hence tomentose; diminutive tomentulose.

torulose: cylindric or nearly so with constrictions or swellings at ± regular intervals.

trailing: growing for some length over the ground, walls or banks.

translucent: transmitting light but not transparent.

transverse: at right angles to the axis.

tree: a woody plant with a distinct trunk or bole and a ± elevated crown.

trichome: any usually hair-like outgrowth of the epidermis.

trigonous: of a solid body triangular in section.

trilete: having 3 scars or fissures on the spore wall, characteristic of tetrahedral pteridophyte spores.

tristylous: having a floral trimorphism in which the anthers and stigmas vary in height among morphs so that the sequence of heights is different in each morph.

truncate: appearing as though cut squarely across, especially a broad square base to a lf.

trunk: the main woody supporting stem of a tree.

tube: a hollow cylinder; diminutive tubule.

tuber: a thickened, usually subterranean, part of a stem functioning as a storage organ; hence tuberous.

tubercle: a small wart-like swelling; hence tubercled, tuberculate, tubercules.

tunic: a loose membrane investing a corm or bulb.

turbinate: top-shaped.

turgid: swollen or fully inflated.

turion: a young sucker or shoot capable of generating a new plant.

twig: a young woody stem, usually the current season's growth of a branch.

type: a specimen to which the name of a taxon at or below the rank of genus is permanently attached.

umbel: an indeterminate infl. with peduncles or pedicels arising from a common centre, and so often umbrella-shaped; hence umbellate, umbelliform; diminutive umbellet.

umbo: a small conical projection from the surface, particularly of gymnosperm cones; hence umbonal.

undulate: waved in a plane at right angles to the surface.

unilabiate:1-lipped.

unisexual: of one sex only.

urceolate: urn-shaped.

utricle: a thin loose cover enveloping some frs.

valvate, valvar: of dehiscing frs, opening by valves; of perianth segments in bud meeting at the edges but not overlapping.

valve: a door-like, often separable, part.

variegated: marked with irregular or regular patches, blotches or lines of 2 or more colours, often the markings lacking chlorophyll and giving the part a diseased appearance.

variety: an infraspecific unit of classification, correctly applied to a regionally distinct part of a sp., often incorrectly used for a cv.

vascular: furnished with specialised conducting tissues.

vein: a strand of conducting and usually strengthening tissue in a lf or similar structure.

velutinous: velvety.

venation: the arrangement of the veins.

ventral: of the inner or lower surface of an organ.

ventricose: of a swelling on the ventral surface.

vernation: the arrangement of lvs in the bud stage.

verrucose: marked with small wart-like excrescences; diminutive verruculose.

versatile: of anthers attached to the filament at or near the middle and able to move freely.

verticel: a whorl of fls or lvs; in Lamiaceae a false whorl of fls that are in cymes; hence verticillate; diminutive verticillasters.

vesicle: a small bladdery sac filled with gas or fluid; hence vesicular.

vestigial: a part now degenerate and non-functional.

vestiture: a covering of any sort, usually hairs or modified hairs.

vexillary: of the broad upper petal in many Fabaceae, especially with reference to the uppermost, often free, stamen.

viable: of spores, seeds, and pollen which is able to germinate.

villous: clad in long soft hairs not matted together.

vine: climbing or trailing woody stemmed plant.

virgate: elongated, straight, slender and pliant, or composed of twigs of this form.

viscid, viscous: sticky.

vittae: the oil tubes embedded in the pericarp of the frs of most Apiaceae.

whipcord: of a flexible branch with appressed, imbricate scale-like lvs.

whorl: an arrangement of 3 or more parts or organs at the same level around an axis.

xeromorphic: of plants adapted to dry conditions.

zygomorphic: having only one plane of symmetry; used especially of irregular fls.

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