Volume II (1970) - Flora of New Zealand Indigenous Tracheophyta - Monocotyledons except Graminae
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Monocotyledones

MONOCOTYLEDONES

KEY TO FAMILIES

Key

1
Plant floating free on fresh water, composed of oval green platelets budding from lateral pouches
Plant attached by roots, differentiated into stem and lvs or bracts
2
2
Plant a high climbing liane
3
Plant not a high climbing liane
4
3
Stem twiner; lvs ± oblong, reticulately veined; infl. laxly racemose
Root climber; lvs long-linear, parallel-veined; infl. a dense spadix
4
Lvs pinnate; infl. axillary, fleshy, enclosed between 2 broad spathes
Lvs simple, sts very reduced; infl. Otherwise
5
5
Ovary clearly inferior below well-developed per.
6
Ovary superior, or per. absent or obscure
9
6
Fls zygomorphic, us. strongly, with one petal differentiated as labellum; central bilaterally symmetric column bearing 1–2 stamens and, on the side facing the labellum, a stigmatic surface
Fls actinomorphic without such column; stamens 6 or 3
7
7
Stamens 3; lvs equitant
Stamens 6; lvs not equitant
8
8
Lvs green; per.-segs free
Lvs represented only by scales, whole plant without chlorophyll; per. tubular
9
Per. petaloid, often coloured at least its inner whorl
10
Per. 0, or membr., glumaceous, or dry, not petaloid
12
10
Lvs oblong to elliptic, spaced along slender stem; petiole short, narrow, twisted to bring smooth convex abaxial lf-surface uppermost
Lvs linear, crowded in tufts; lf-base sheathing, us. broad
11
11
Stem stout, woody, sts forming heavy trunk with terminal lf-tufts; lvs very fibrous
Stem stout to slender, us. completely hidden within lf-bases; lvs us. not strongly fibrous
12
Fls unisexual in capitula on simple or branched infls; capitula all globose, lower ones female, upper male
Fls bisexual, or if unisexual infls otherwise
13
13
Per.-segs us. not obscured by bracts though mostly small; fls bisexual
14
Per.-segs 0, or plumose, or microscopic and not easily visible; fls uni- or bi-sexual
16
14
Per.-segs 4; lvs mostly broad, if linear wholly aquatic and permanently submerged
Per.-segs 6; lvs linear to terete or reduced to bladeless sheaths
15
15
Infl. an ebracteate raceme or spike
Infl. a ∞-fld cyme, much-branched to condensed, or fls sts solitary; floral bracts present
16
Fls in true or apparent (sts 1-fld) spikelets, each fl. us. subtended by membr. bract(s)
17
Fls otherwise arranged, sts solitary
20
17
Lvs reduced to ∞ sheathing bracts along stem; plants dioec
Lvs with well-developed lamina, or if reduced to sheathing bracts these few and mostly basal; plants rarely dioec
18
18
Lvs ± jointed at junction of lamina with sheath, us. with definite projecting ligule; spikelets distichous; each fl. associated with 2 bracts, one below, one above
23. GRAMINEAE
Lvs not jointed at junction of lamina with sheath; ligule if present not projecting; spikelets distichous or cylindric; each fl. in axil of a single bract, or occ. ebracteate
19
19
Fls grouped in simple spikelets; each fl. subtended by a bract (glume) which is visible without dissection of infl.
Fls grouped in a complex, ± spike-like infl. within 2–3 glume-like bracts; individual fls either without subtending bracts or enclosed in a hyaline scale only visible by dissection
20
Lvs standing freely erect; infls large, cylindric, dense
Lvs flaccid and incapable of standing erect without support of water; infls small and inconspicuous
21
21
Fls borne on a flattened axis permanently enclosed in a spathe-like lf-sheath; marine, intertidal to subtidal
Fls borne on a cylindric axis breaking out of a short lf-sheath; brackish to fresh water
22
22
Infl. a 2-fld spike; ripened carpels long-stalked, ± umbellately arranged at top of long spirally twisted peduncle
Infl. of 1-several fls; frs short-stalked, apparently axillary, solitary or in small clusters

KEY TO GENERA

(Gramineae appear only as a Family. For a full key to Orchidaceae see p. 104; only a few genera of orchids come out separately here.)

Key

1
Lvs with well developed laminae, sts stem-like in rushes and sedges
2
Lvs either 0 or reduced to scales or sheathing bracts
6
2
Lvs pinnately compound
GROUP A RHOPALOSTYLIS
Lvs simple
3
3
Lf bearing hairs, short to long and sts inconspicuous; or lf wholly or partly covered with appressed scales
GROUP B 7
Lf glab. but margins and nerves may be scabrid
4
4
Per. petaloid, us. white or coloured
GROUP C 15
Per. not petaloid, us. glumaceous, membr., of bristles, or 0
5
5
Lf-margins not at all scabrid
GROUP D 26
Lf-margins scabrid to the touch, either throughout or near tip only
GROUP E 45
6
Plant with chlorophyll; stems green, sts with admixture of tawny colours
GROUP F 57
Plant without chlorophyll; stems mostly white or dull brown, never green
GROUP G 69

GROUP B

Lvs simple, bearing short or long hairs, or appressed scales

Key

7
Lvs bearing ± terete hairs but no flat scales
8
Lvs bearing scales, us. longest on sheaths, sts associated with felted hairs; scales mostly elongated and flakelike, sts fused into a silvery detachable pellicle
14
8
Plant an epiphyte; each lf seated on a globose green pseudobulb
Plant terrestrial; lvs not associated with green pseudobulbs
9
9
Plant perennating by subterranean tubers; lvs dying off annually; per. petaloid
10
Plant evergreen or strictly annual; per. either not petaloid or 0
11
11
Per. of 6 chaffy scales; fr. a 3-seeded capsule
Per. either 0 or represented only by 2 minute scales; fr. 1-seeded
12
12
Stems hollow; spikelets distichous and very rarely solitary, sts 1-fld
GRAMINEAE
Stems solid; infl. never a true distichous spikelet, but if apparently so then solitary
13
13
Fls in true spikelets; glumes ∞ and spirally arranged
Fls in a solitary, complex but superficially spikelike infl. enclosed within 2 almost opp. glumelike bracts
14
Tepals all membr., connate below into a cup-shaped tube; lf-sheath not keeled, crescentic in T.S.; plants too large to be turf-forming
Tepals either fleshy or, if membr., free almost to base; lf-sheath us. keeled and ± V-shaped in T.S. except in very small turf-forming spp.

GROUP C

Lvs simple with well-developed laminae, glab., but margins and nerves may be scabrid; per. petaloid

Key

15
Fr. a capsule
16
Fr. a berry
24
16
Fls zygomorphic, us. strongly; ovary 1-locular, inferior
Fls almost to quite actinomorphic; ovary 3-locular, mostly superior
17
17
Lvs laterally flattened and iris-like throughout their length, equitant at base
18
Lvs dorsiventrally flattened; if equitant at base lamina is keeled in upper part
19
18
Ovary inferior; fls white
Ovary superior; fls red in all parts
19
Staminal filaments ornamented with hairs; infl. us. an openly branched panicle, if reduced and ± raceme-like then an occ. bract subtends 2 fls.
Staminal filaments glab.; infl. us. truly racemose or of only 1–2 fls, if paniculate then narrowly so and almost woody
20
20
Lvs all dying down at end of season; plant perennating by swollen underground parts; fls never dull red or blue
21
Lvs green throughout year; plant lacking subterranean storage organs; fls never clear yellow
23
21
Ovules 2 per locule; seeds as long as capsule; storage organs ∞ fleshy roots
Ovules ∞ per locule; seeds much shorter than capsule; storage organ a corm
22
22
Ovary inferior, wall breaking irregularly to release seeds; fls yellow; corm subglobose, enclosed in reticulate fibres
Ovary superior; capsule of 1–3 valves, opening widely; fls white or pinkish; corm ± pear-shaped, enclosed in membr. sheaths
23
Plant resembling a small rhizomatous grass; per. spread out almost flat, blue to white
Plant composed of crowded fans of large, excessively fibrous lvs; per. narrow-elongated, greenish to dull red
24
Lvs ± oblong, not sheathing at base, spaced out singly along wiry stems
Lvs linear, sheathing at base, grouped in tufts
25
25
Stem scarcely woody, almost entirely enclosed in lvs; midrib scabrid abaxially
Stem woody, sts forming long trunk; midrib not scabrid abaxially

GROUP D

Lf simple, sts stem-like, glab., margin not scabrid; per. either not petaloid, or absent

Key

26
Plant a woody liane
Plant of a different habit
27
27
Aquatic plants normally entirely submerged or inter-tidal; lvs mostly filamentous to ribbon-like, us. ± translucent
28
Land plants or aquatics with floating lvs, or stems or lvs raised above the water; lvs if narrow not translucent
33
28
Plants of sites below regular high tide mark, us. forming grass-like sward; lvs linear but not filiform, with ∞ transverse veinlets at right angles to midrib and strong marginal nerves
Plants of brackish or fresh water; lvs us. distributed on slender stems, or, if closely crowded, almost filamentous, without ∞ transverse veinlets and strong marginal nerves
29
29
Peduncles long, threadlike, becoming spirally coiled as fr. matures; fls 2, sessile and close together, each producing (2)–4–8–(16) obviously stalked achenes
Peduncles mostly short, never spirally coiled; fls 1-∞, each producing at most 4 sessile or obscurely stalked achenes
30
30
Lvs without sheaths or stipules, tufted, all radical; plant rarely as much as 5cm. tall
Lvs with stipules, sts sheathing; stem-internodes mostly easily visible, many > 1 cm. long
31
31
Fls bisexual, tetramerous, ∞, in obvious, fully exserted, continuous or interrupted spikes.
Fls unisexual, not tetramerous, solitary or few in inconspicuous clusters
32
32
Lvs in false whorls of 3–4; stipules free
Lvs obviously alternate, mostly distant; stipules adnate to lf-base
33
Fr. a capsule; seeds ∞ per carpel
34
Fr. not a capsule; seeds 1 per carpel
36
34
Infl. of several to ∞ fls
Infl. of one solitary fl.
35
35
Floral bracts = or > tepals
Floral bracts much < tepals
36
Lvs elliptic to broadly ovate
Lvs of linear order
37
37
Per. small but easily discernible
38
Per. either 0 or microscopic and ± hidden
39
38
Fls in compact, globose, obviously unisexual heads
Fls bisexual in narrow raceme
39
Fls and associated structures all threadlike, densely crowded in large cylindric velvety-brown infls; lvs ± plano-convex in T.S., spongy within
Fls and associated bracts and glumes not wholly threadlike; infls if cylindric not with fine velvet-textured surface; lvs not as above
40
40
Lvs mostly linear with marginal sclerenchyma, us. keeled and ligulate
GRAMINEAE
Lvs mostly linear-filiform to terete or trigonous, a few equitant and iris-like, almost all eligulate
41
41
Fls in many-glumed spikelets
42
Fls in a solitary, complex but superficially spike-like infl. enclosed within 2–3 glume-like floral bracts
44
42
Glumes spirally arranged
Glumes distichous
43
43
Lvs either ± horizontal with narrow flat lamina or erect and ± terete but not septate
Lvs erect, either with broad flat lamina, or terete with septa visible externally or internally
43A
43A
Fr. with hard bony endocarp and ± spongy, thick or thin mesocarp, sessile or scarcely stipitate
Fr. with thin brittle pericarp, distinctly stipitate
44
Floral bracts 2, appearing opp.; lf-sheaths hyaline, pale green or colourless
Floral bracts (2)–3, each clearly overtopping the one below; lf-sheaths opaque, light brown

GROUP E

Lvs simple, sts stem-like; margins scabrid throughout or near tip only; per. not petaloid

Key

45
Plant a root-climber attaching itself by coarse aerial roots produced from thick, hard but brittle stem
Plant not a root-climber; aerial roots, if any, contributing to thickness of "trunk"
46
46
Lf ± jointed at junction of lamina with sheath, us. with definite projecting ligule
GRAMINEAE
Lf not jointed at junction of lamina with sheath, ligule, if any, not projecting
47
47
Fr. enclosed in a utricle
48
Fr. not enclosed in a utricle
49
48
Rhachilla produced into a hook
Rhachilla not produced into a hook
49
Lvs harshly scabrid, almost razor-sharp, on margins throughout and on nerves
50
Lvs scabrid, but not lacerating, on margins throughout or only towards lf-tip
53
50
Spikelets ∞-fld
51
Spikelets 1–2-fld
52
51
Glumes distichous; compound infl. broad; subinfls often stalked, ± radiately arranged
Glumes spiral; compound infls long; subinfls sessile and mostly confluent, ± spirally arranged
52
Culms terete; glumes spirally arranged
Culms trigonous; glumes distichous
53
Per. of 6 minute hypog. scales
54
Per. either 0 or of hypog. bristles
55
54
Lvs similar to stems; hypog. scales attached to nut and falling with it; plants erect, of coarse tufted habit
Lvs unlike stems; hypog. scales remaining attached to stem after nut falls; plants low-growing, forming dense or loose cushions
55
Glumes spirally arranged
Glumes distichous
56
56
Per. of 6 setae, plumose throughout
Per. either 0 or of 6–1 setae, non-plumose, or plumose at base only

GROUP F

Lvs 0 or reduced to sheaths; plant with chlorophyll

Key

57
Plants firmly rooted in soil; stems terete or tetragonous, ± erect
58
Plants unattached, us. free-floating on surface of fresh water; each plant consisting of a single oval green platelet or a few budding laterally from one another
67
58
Bract-like sheaths ∞, spaced at intervals along stem
59
Bract-like sheaths all or most crowded at stem-base
61
59
Stems unbranched
Stems branched
60
60
Infl. a panicle
Infl. of distant spikelets towards stem-tip
61
Per. of 6 chaffy tepals
Per. either 0, or of minute scales or bristles
62
62
Per. of 6 minute scales attached to base of nut
Per. either 0, or of bristles
63
63
Glumes spirally arranged
64
Glumes distichous
65
64
Style swollen at base
Style not swollen at base
65
Bract-like sheaths dark red-purple, almost black; rhachilla of spikelet flexuous; style-base not persistent on nut
Bract-like sheaths light brown or reddish; rhachilla of spikelet straight; style-base ± pubescent, persistent on nut
66
66
Persistent pubescent style-base merging into top of nut, never more than ½ length of nut, often much shorter
Persistent pubescent style-base distinct at top of nut, narrower than but almost as long as nut
67
Roots absent; platelets deeply keeled
Roots present; platelets flat to slightly bi-convex, without deep keel
68
68
Roots 1 per platelet
Roots 2 or more per platelet

GROUP G

Lvs 0 or reduced to scales or sheathing bracts; plants without chlorophyll

Key

69
Erect stem tall, carrying raceme of fls well above soil-surface
70
Erect stem very short, fls solitary, ± hidden in lf-mould
71
70
Per.-segs connate into tube; subterranean tubers swollen and fusiform
Per.-segs free; subterranean rhizomes elongated and uniformly narrow throughout
71
Fls actinomorphic, lantern-shaped, mostly red
Fls strongly zygomorphic, with large frilled labellum, white, at most flecked with red
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