Volume I (1961) - Flora of New Zealand Indigenous Tracheophyta - Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledons
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Forstera "flagriformes"

F. "FLAGRIFORMES"

Small shrubs with terete or tetragonous whipcord-like branches covered with scalelike lvs. Internodes bifariously white-hairy, photosynthetic and very similar in appearance to lvs, us. short and almost entirely hidden; nodal joint at base of lf not always externally recognizable. Lvs sessile, connate, ± appressed but not adnate, us. overlapping, concavo-convex, often keeled, sts ribbed, mostly thickly fleshy when fresh, margined with fine white cilia, conspicuously dotted with stomata on abaxial surface. Fls sessile in simple, short, crowded terminal spikes continuous with branchlets; bracts opp., similar to lvs but us. larger and thinner; rhachis and bases of bracts and calyx clad in abundant white woolly hairs (except in H. c upressoides). Calyx-lobes ciliolate; anterior pair flat or slightly convex, fused in lowest third or for whole length, overlapping posterior lobes at deep lateral clefts; posterior lobes often smaller, keeled, separated by wide deep cleft or sts by rudimentary fifth lobe, fused in H. cupressoides. Corolla us. white, lobes 4, unequal, spreading, = or > tube. Capsule glab., dorsally compressed with septum across broadest diam. (except in H. cupressoides), laterally grooved near apex, dehiscing septicidally with only slight loculicidal split. Fourteen spp., from upland and mountain habitats.

Key

1
Plant a dense, rounded, ± glaucous bush with erect branchlets not > 1 mm. diam. and remote lf-pairs; capsule laterally compressed and ± didymous, scarcely 2 × 1 mm
Plant not at all as above; capsule dorsally compressed, not didymous, much larger
2
2
Nodal joint well-marked, though often hidden by overlap of lvs; anterior calyx-lobes us. only ⅓ fused (completely fused in H. armstrongii)
3
Nodal joint obscure, though us. exposed, the lf appearing ± continuous with internode below; anterior calyx-lobes almost completely fused
12
3
Lvs ribbed or striped on outer surface, at least for short distance from upper margin
4
Lvs sts wrinkled when dry but not regularly ribbed or striped
6
4
Lvs abruptly narrowed to thickened cusp or spine at apex or at least with thickened acute tip, us. ribbed only near upper margins, sts rather faintly
Lvs with broadly subacute to acute, rounded or ± keeled tip, strongly ribbed for most of length
5
5
Plant forming small tufts 5-15 cm. tall; branchlets 1·5-2 mm. diam
Plant robust, 40-60 cm. tall; branchlets 2-2.5 mm. diam.
6
Branchlets strongly tetragonous, 2-3 mm. diam.; lvs rarely < 2.5 mm. long, ± deltoid with long, keeled, thickened tip us. reaching beyond connate margins of lf-pair above
Branchlets terete or, if weakly tetragonous, not > c. 2 mm. diam.; lvs not > 2.5 mm. long, tip not elongated, not or hardly reaching beyond connate margins of lf-pair above
7
7
Lvs 1-1·5 mm. long, internodes visible for at least half lf-length
8
Lvs rarely < 1·5 mm. long, internodes hidden or visible for < half lf-length
9
8
Lvs rounded and incurved at apex
Lvs with small sharp tip
9
Branchlets terete, rarely < 2.5 mm. diam.; lf-apex smoothly appressed or convexly rounded, obtuse or subacute or with abrupt, very small, blunt mucro
Branchlets ± terete to weakly tetragonous, rarely > 2 mm. diam.; lf-apex convexly rounded or slightly keeled, obtuse to acute but never mucronate
10
10
Lf-tip subacute, convexly incurved and thickened to be ± boat-shaped; nodal joints us. hidden; corolla-tube us. < calyx (plant of North Id)
Lf-tip obtuse to acute, sts slightly convexly incurved but hardly boat-shaped; nodal joint us. visible; corolla-tube = or > calyx (plant not of North Id)
11
11
Branchlets ± tetragonous, 2-2.5 mm. diam., rarely glossy; branches stout, erect or procumbent, often naked below with distal tufts of close-set, spreading branchlets 2-3 cm. long (plant of Fiordland and Stewart Id)
Branchlets not or scarcely tetragonous, 1·5-2 mm. diam., us. glossy; branches slender, us. arching and bearing flexible branchlets up to 5 cm. long on upper side for most of length (plant of Nelson)
12
Lvs thick, keeled; branchlets us. glossy, firm in texture and ochre-coloured when fresh
Lvs thin, not keeled; branchlets not glossy, flexible or succulent and green when fresh
13
13
Lvs with small sharp tip, widely spreading and encircling branchlet when dry
Lvs obtusely rounded or truncate, only slightly spreading when dry
14
14
Plant low and spreading with profuse mass of small branchlets; internodes 1-1·5 mm. long, lvs us. slightly overlapping
Plant erect, more sparingly branched; internodes 2-3 mm. long, lvs not at all overlapping

The so-called "whipcord" hebes are well-defined by their peculiar habit, though they are quite closely related to the "Buxifoliatae". Some of the spp. are very difficult of discrimination and it has not been possible to construct a key that will always provide a clear separation. Hybrids between whipcord spp. have not certainly been detected but crossing not uncommonly occurs with other spp., particularly the "Buxifoliatae", and some hybrids are well-known in cultivation.

Measurements given of the diameter of ultimate branchlets are taken from below the extreme tip and include the lvs. In spp. with very close-set overlapping lvs internode-length is most easily found by halving the distance between the apices of alt. lf-pairs. In those spp. where the position of the node is not externally obvious it may be determined by breaking off the lf. Plants alter considerably on drying, lvs which were smoothly appressed shrinking and ± standing out from the branchlets and internodes sts longitudinally wrinkling. This is particularly noticeable in specimens from cultivated plants, which also us. have unnaturally long internodes and often revert to semi-juvenile foliage. Juvenile and reversion lvs have not been seen in all spp. but in all cases described are spreading, petiolate, membr., entire or pinnatifidly toothed or lobed, passing by gradual transition into adult form. Fls sts appear as early as November or December, but for most spp. the main flowering appears to be in January or February with late fls until April and May.

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