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Volume V (2000) - Flora of New Zealand Gramineae
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Chionochloa Zotov

Chionochloa Zotov, 1963

Type species: C. rigida (Raoul) Zotov

Perennial tussocks, solitary or occasionally sward forming; branching intravaginal or sometimes extravaginal. Leaf-sheath long persistent and clothing shoots, entire or fracturing into irregular segments, glabrous or with internerve hairs, tuft of hairs at apex. Ligule a ring of hairs usually c. 1 mm. Leaf-blade 5 cm to 1.5 m long, glaucous, flat to V-shaped, or junceous, persistent or disarticulating at ligule or falling with part of sheath, abaxially usually glabrous, occasionally hairy, adaxially prickle-toothed or papillate, rarely unornamented, margins usually long hairy below. Culm to 2 m, erect, exceeding leaves. Inflorescence an open or compact panicle of few to many spikelets, glabrous to conspicuously long hairy, usually with long hairs at branch axils. Spikelets of few to several ⚥ florets, disarticulating above glumes and between florets. Glumes unequal, linear, acute or rarely awned, usually glabrous; lower 1-3-nerved, upper 3-5-7-nerved. Lemma (7)-9-nerved, nerves anastomosing below sinus, lateral lobes awned or triangular-acute, densely hairy on margins and in columns in all, few, or no internerves; central awn straight or reflexed from twisting, flat or indistinct column from sinus. Palea > lemma sinus, keels ciliate, interkeel infrequently prickle-toothed, flanks long hairy below. Callus short, blunt, hairs long covering lemma; disarticulation oblique. Rachilla usually glabrous. Lodicules 2, irregularly rhomboidal and lobed, nerved, margins long hairy especially apically. Stamens 3; anthers caudate, shorter in male-sterile flowers. Gynoecium bistylar, ovary glabrous, < lodicules. Caryopsis free, obovate, smooth or rugose; embryo ⅓-½ caryopsis length; hilum linear, ½-⅔ caryopsis length. Fig. 16.

SYNOPSIS

  • A.
    • Leaf-blade persistent on sheath
    • 1.
      • Lemma scabrid with prickle-teeth
      • (a)
        • Leaf-blade margin smooth
        • (i)
          • Mat-forming; alpine:
          • 3. australis
        • (ii)
          • Caespitose; coastal cliffs:
          • 4. beddiei, 5. bromoides
      • (b)
        • Leaf-blade margin scabrid:
        • 6. cheesemanii, 11. flavicans
    • 2.
      • Lemma lacking prickle-teeth
      • (a)
        • Leaf-blade gramineous
        • (i)
          • Sheath long, margin flat:
          • 14. macra, 17. pallens
        • (ii)
          • Sheath short, margin undulating:
          • 15. oreophila
      • (b)
        • Leaf-blade junceous:
        • 12. juncea, 19. rubra subsp. cuprea
  • B.
    • Leaf-blade disarticulating at ligule
    • 1.
      • Leaf-blade gramineous
      • (a)
        • Leaf-blade margin glabrous
        • (i)
          • Leaf-blade tip blunt:
          • 2. antarctica, 13. lanea
        • (ii)
          • Leaf-blade tip pungent:
          • 16. ovata
      • (b)
        • Leaf-blade margin hairy below
        • (i)
        • (ii)
          • Inflorescence glabrous:
          • 17. pallens subsp. cadens, 22. vireta
    • 2.
  • C.
    • Leaf-blade falling with part of sheath below ligule
    • 1.
      • Leaf-blade gramineous:
      • 10. flavescens, 18. rigida
    • 2.
      • Leaf-blade junceous
      • (a)
        • Leaf-sheath strict:
        • 9. defracta, 19. rubra
      • (b)

Key

1
Leaf-blade junceous
2
Leaf-blade gramineous
9
2
Leaf-blade disarticulating at ligule or falling with part of sheath
3
Leaf-blade persistent on sheath
7
3
Leaf-blade falling with part of sheath
4
Leaf-blade disarticulating at ligule
6
4
Abaxial leaf-blade evidently hairy; inflorescence hairy
Abaxial leaf-blade glabrous; inflorescence glabrous
5
5
Leaf-sheath erect or strict; adaxial leaf-blade with prickle-teeth or papillae, margin scabrid or smooth
5a
Leaf-sheath entire
Leaf-sheath fracturing into short segments
5b
5b
Adaxial leaf-blade with short hairs above ligule or 0
Adaxial leaf-blade with weft of long hairs above ligule
Leaf-sheath coiling up; adaxial leaf-blade with prickle-teeth, margin smooth
6
Abaxial leaf-blade with two columns of antrorse hairs
Abaxial leaf-blade glabrous
7
Leaf-sheath glabrous, margin undulating, long entangling hairs at apex; leaf-blade to 10 cm
Leaf-sheath hairy, margin often coiling up; leaf-blade to 1 m
8
8
Adaxial leaf-blade with long hairs at midpoint; margin with long hairs below and prickle-teeth above
Adaxial leaf-blade with prickle-teeth at midpoint; margin with long hairs below, becoming glabrous
9
Leaf-blade persistent on sheath
10
Leaf-blade disarticulating at ligule or falling with part of sheath
16
10
Adaxial leaf-blade scabrid at midpoint
11
Adaxial leaf-blade glabrous at midpoint
13
11
Leaf-sheath to 5 cm, thin, margins wavy; mat-forming
Leaf-sheath 15-20 cm, firm, margins flat; caespitose
12
12
Leaf-sheath keeled; leaf-blade V-shaped
12a
Leaf-blade disarticulating at ligule
Leaf-blade persistent on sheath
12b
12b
Adaxial leaf-blade with prickle-teeth or papillae
Adaxial leaf-blade with long hairs and prickle-teeth or papillae
Leaf-sheath rounded; leaf-blade flat to shallowly U-shaped
13
Leaf margin scabrid
14
Leaf margin smooth
15
14
Branching intravaginal; sheath densely hairy
Branching extravaginal; sheath glabrous
15
Adaxial leaf-blade with weft of short hairs above ligule
Adaxial leaf-blade with weft of long hairs above ligule
16
Adaxial leaf-blade with weft of hairs above ligule or with interlocking long hairs
17
Adaxial leaf-blade lacking weft of hairs or interlocking hairs
20
17
Leaf-sheaths persistent and entire
18
Leaf-sheaths breaking into short segments
19
18
Abaxial leaf-blade very scabrid above; sheath compressed and flat; innovations extravaginal (North, South and Stewart Is)
18a
Leaf-sheath conspicuously long hairy; adaxial leaf-blade with weft of long hairs at base
Leaf-sheath and adaxial leaf-blade glabrous
Abaxial leaf-blade smooth above; sheath terete; innovations intravaginal (Auckland and Campbell Is)
19
Leaf-blade to 10 mm wide, tapering, often reddened above; sheath to 20 cm, coloured or pale; shoots to 10 mm diam., stout
19a
Leaf-blade with weft of long hairs above ligule
19b
Leaf-blade with weft of short hairs or lacking hairs above ligule
19c
19b
Leaf-sheath glabrous except on margin
Leaf-sheath hairy
19c
Leaf-blade lacking hairs above ligule
Leaf-blade with weft of short hairs above ligule
Leaf-blade to 7 mm wide, narrowing, often golden above; bronze sheath to 30 cm; shoots to 5 mm diam., slender
19d
Inflorescence glabrous except for long hairs at branch axils; leaf-blade margin hairy below becoming scabrid
Inflorescence hairy or scabrid; leaf-blade margin becoming smooth below
20
Leaf-blade margin smooth above, long hairs below
21
Leaf-blade margin prickle-toothed above, long hairs below
23
21
Adaxial leaf-blade papillate; sheath stout and stiff
21a
Leaf-blade curved, coriaceous, tapering abruptly to a very pungent point (Stewart Id)
Leaf-blade curved or twisting above, ± flexible, tapering gradually to a long, thin, pungent apex (South Id)
21b
21b
Shoots from short stout stems; leaf-blade stout, to 6 mm wide, curled or twisting above
Shoots from very long prostrate stems; leaf-blade slender, to 3 mm wide, straight or somewhat twisting above
Adaxial leaf-blade prickle-toothed
22
22
Ligule to 3.5 mm; sheath becoming fibrous, short hairy
Ligule to 1 mm; sheath intact, very long (4 mm) hairy
23
Plants slender; sheath margins thin and undulating; leaf-blade ± flat, to 2.5 mm wide
Plants stout; sheath margins firm and flat; leaf-blade V-shaped, to 10 mm wide

24 spp. of Australasia. Endemic spp. 22.

N.Z. spp. were revised by Connor, H. E. N.Z. J. Bot. 29: 219-283 (1991), and synoptically treated by Zotov, V. D. N.Z. J. Bot. 1: 78-136 (1963). They are collectively known as snow-tussock or snow-grass except for a few.

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