Volume IV (1988) - Flora of New Zealand Naturalised Pteridophytes, Gymnosperms, Dicotyledons
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Astereae Cass.

ASTEREAE Cass.

Annual or perennial herbs or shrubs, rarely small trees, not usually strongly scented, lacking latex. Lvs usually alternate or basal, very rarely opposite. Involucral bracts in (2)-several rows, herbaceous to chartaceous, rarely membranous at margins or apex. Receptacle usually without scales, very rarely with scales. Capitula usually ⚥, rarely unisexual, sometimes homomorphic-discoid, but usually heteromorphic with 1-several rows of outer, filiform to ligulate, ♀ florets, and few to numerous, actinomorphic, ⚥ or rarely ♂ disc florets. Anther bases usually obtuse, rarely tailed. Style branches each with 2 stigmatic lines. Achenes usually homomorphic, rarely heteromorphic, with 0-few ribs or wings, terete or compressed; pappus usually of long scabrid to plumose hairs, sometimes of short hairs, scales or barbed awns, or rarely reduced to a corona or 0.

Key

1
Pappus 0, or reduced to short bristles or rigid barbed awns
2
Pappus of disc and usually ray florets of long scabrid to plumose hairs, sometimes with an outer row of shorter hairs or scales
7
2
Ligules yellow; pappus of rigid barbed awns
Ligules white, flushed pink or lilac, or 0; pappus 0, or of short, simple bristles
3
3
Achenes distinctly beaked at apex
Achenes rounded at apex
4
4
Ligules 0
5
Ligules conspicuous
6
5
Perennial; lvs mostly basal; achenes narrowly ribbed
Annual; lvs mostly cauline; achenes with dissected marginal wings
6
Peduncle with simple antrorse hairs; pappus 0
Peduncle glabrous or with glandular hairs; pappus usually of short bristles, rarely 0 in mature achenes
BRACHYCOME
7
Plant a hard-wooded shrub or tree
8
Plant a herb, at most woody toward base
11
8
Dioecious; ♂ capitula with all florets tubular; ♀ capitula with all florets filiform
Capitula with outer florets ♀ and filiform to ligulate, and inner florets ⚥ and tubular, or all florets ⚥ and tubular
9
9
Plant low-growing with prostrate rambling branches
Plant an erect or ascending shrub or tree
10
10
Longest involucral bracts < 13 mm; capitula solitary or in compound infls; pappus hairs unequal
OLEARIA†
Longest involucral bracts > 14 mm; capitula solitary; pappus hairs equal
11
Lvs all or mostly in tufts or rosettes at ground level
12
Lvs all or mostly scattered and cauline
14
12
Capitula in racemes
Capitula solitary
13
13
Disc corollas yellow; pappus scabrid
Disc corollas purple; pappus plumose
14
Ligules yellow
Ligules white, blue, red, pink, or violet to purple
15
15
Ligules 0 or inconspicuous, at most ± = pappus
16
Ligules conspicuous, exceeding pappus
17
16
Plant usually completely glabrous; bracts tinged purple at apex and margins
Plant with at least a few hairs on upper stem or upper surface of lvs, if almost glabrous then bracts with a distinct purple spot at apex
17
Involucral bracts distinctly dimorphic, the outer herbaceous, the inner completely membranous; annual
Involucral bracts all similar, but the inner usually more chartaceous than the outer; annual or perennial
18
18
Perennial herbs with erect stems arising annually from rhizomatous crown
Annual or short-lived perennial erect herbs, or procumbent to erect somewhat woody herbs with perennial stems lacking rhizomes
19
19
Pappus of 2-several rows of long hairs
Pappus of 1 row of long hairs, sometimes with a short outer row of scales and the long hairs lacking on the ray achenes

135 genera, c. 2500 spp., cosmopolitan but mainly temperate, especially Africa and America.

This large tribe includes many genera cultivated for their showy capitula. As in other tribes, the limits of large genera are not well defined and authors differ in the number of segregate genera they accept. The treatment here generally follows that of Grau, J., in Heywood et al. (op. cit.).

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