Volume V (2000) - Flora of New Zealand Gramineae
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Bambuseae

BAMBUSEAE *

by W.R. Sykes

Evergreen grasses, usually with woody culms (bamboos), sometimes herbaceous (not in N.Z.), occasionally climbing. Growth either sympodial (determinate) with short and thick rhizomes (pachymorph) which form dense clumps or monopodial (indeterminate) with short to long ± slender rhizomes (leptomorph) which form ± diffuse thickets. Culm nodes marked by 1 or 2 prominent rings, the lower representing the scar from a modified sheathing leaf (culm-sheath), internodes usually hollow. Culm-sheath with sheath proper, membranous ligule and often auricles and associated oral bristles, and a short, thickened, sometimes caducous sheath-blade. Branches 1 to many, often only from middle and upper nodes (nodes often many-branched because of rebranching at base from the single bud). Leaf-sheath with ligules and often auricles and oral bristles. Leaf-blade petiolate, often tessellate (cross veins forming a lattice). Inflorescence of panicles, racemes, or spikes often subtended by a leaf-like spathe; sometimes a condensed fascicle with branches very contracted and small bracts subtending the spikelets. Spikelets all similar, 1-many-flowered. Glumes (0)-2-(4) (florets often Ø in countries of introduction). Palea exposed or ± enfolded by lemma. Lodicules usually 3. Stamens 3-6 (in N.Z.). Gynoecium: style 1; stigmas 1-3. Fruit usually a caryopsis; hilum linear.

Key

1
Culm internodes flattened or grooved on alternate sides or obtusely quadrangular, at least in upper culm
2
Culm internodes terete, occasionally with a few uppermost internodes flattened on alternate sides
4
2
Culm internodes above lowest branches obtusely quadrangular; basal nodes with short hard or spine-like adventitious rootlets; branches 3 at each node, subequal
Culm internodes above lowest and middle (or sometimes only upper) branches flattened and/or grooved on alternate sides; basal nodes lacking spine-like adventitious rootlets; branches 2-(3) at each node, unequal (excluding rudimentary branch if present), later sometimes clustered
3
3
Culm internodes flattened or grooved on alternate sides throughout or to lowest branch-bearing node; nodes with a 2-branched system which appears diffuse
Culm internodes terete, some flattened or grooved on alternate sides; nodes ultimately with up to 8 ± dense branches
4
Middle culm nodes usually with one branch; culm-sheaths persistent; rhizomes monopodial
5
Middle culm nodes with several to many branches; culm-sheaths deciduous, sometimes tardily so; rhizomes sympodial or monopodial
7
5
Culm 3-5 m; lower and middle culm-sheaths > internodes; oral bristles smooth, or 0
Culm < 2 m (in wild and cultivated N.Z. plants); lower and middle culm-sheaths < internodes; oral bristles scabrid throughout or below, rarely 0
6
6
Culms ascending; oral bristles scabrid along their whole length
Culms strictly erect; oral bristles scabrid only near base
7
Culms > 3 m × > 1 cm diam. (often very much higher and thicker); new culms appearing in winter; leaf-blade not tessellate; inflorescence a dense or capitate spike
Culms usually < 3 m × < 1 cm diam.; new culms appearing in late spring and summer; leaf-blade generally tessellate; inflorescence a diffuse panicle
8
8
Rhizomes running, sometimes shortly so, branching always monopodial; plant dense but not caespitose; culm-sheaths ± persistent; leaf-blade obviously tessellate
Rhizomes very short, branching sympodial; plant densely caespitose; culm-sheaths deciduous; leaf-blade not obviously tessellate

Since flowering is so irregular, bamboos are usually identified by vegetative features. As far as possible the generic descriptions below are comprehensive for species naturalised in N.Z. Other genera are in cultivation here; a few of these are mentioned because they have been sometimes confused with taxa treated as naturalised.

Flowering in bamboos is usually infrequent and irregular, often many years elapsing between flowering periods, although some species flower annually. Synchronous flowering is a feature of many species, following which all the plants may die or, as with most species in N.Z., parts of the clump die. Sometimes this happens for several years in succession. However some species have never flowered in this country since they were introduced, and some have probably never flowered anywhere in cultivation.

The treatment of Bambuseae here includes genera accepted by Stapleton, C. Bamboos of Nepal: an illustrated guide (1994), and by McClintock, D. in Walters, S. M. et al. (Eds) European Garden Flora Vol. II (1984).

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