Volume IV (1988) - Flora of New Zealand Naturalised Pteridophytes, Gymnosperms, Dicotyledons
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Helichrysum Mill.

HELICHRYSUM L.

Usually herbs or shrubs, rarely lianes. Lvs usually alternate, rarely subopposite, simple, entire. Capitula solitary or in small corymbs; corymbs sometimes aggregated into dense or diffuse clusters or panicles. Involucral bracts in 2-many rows, imbricate, membranous, sometimes with coloured radiating lamina. Receptacle flat to convex; scales usually 0, rarely few and reduced. Outer florets ♀, filiform, usually few or sometimes 0. Inner florets (sometimes all florets) ⚥, tubular. Achenes all similar, terete or slightly flattened, obscurely angled or ribbed; pappus hairs in 1 row, narrow or thickened at apex.

Key

1
Lvs spreading, distant
2
Lvs of mature plant appressed to stem, imbricate
7
2
Erect, annual herb; capitula > 25 mm diam.
Creeping perennial herb, or woody liane or shrub; capitula < 25 mm diam.
3
3
Capitula 5-numerous in clusters or cymes
4
Capitula solitary
5
4
Lvs apetiolate, with paper-thin grey tomentum on upper surface
Lvs appearing distinctly petiolate, glabrous on upper surface when mature
5
Scandent shrub or liane; lvs of 2 forms - some spreading, some appressed and imbricate
Creeping herb; lvs all spreading
6
6
Middle and inner involucral bracts with conspicuous milky white (rarely rose or cream), radiating lamina; lvs around whole stem
Involucral bracts inconspicuous, pale brown, not radiating; lvs in 2 rows
7
Stems with exposed green lf apices and white tomentum
8
Stems usually appearing grey-, brown-, or yellow-tomentose, if white then lf apices not exposed
10
8
Stems (4)-5-10 mm diam. (including lvs); lvs (4)-5-6 × 2-3 mm
Stems 1-4 mm diam. (including lvs); lvs 2-4 × 0.3-2 mm
9
9
Stems 1-2 mm diam. (including lvs); lvs 0.3-1 mm wide; achenes densely hairy
Stems 2-4 mm diam. (including lvs); lvs 1-2 mm wide; achenes with short appressed hairs
10
Stems brown, usually with lvs spreading on part of stem
Stems usually yellowish or grey, rarely white, with all lvs appressed and imbricate in mature plants
11
11
Stems usually yellowish, rarely white, 2-4 mm diam. (including lvs); outer surface of lf tomentose, but finally partly glabrous
Stems grey, 1-2 mm diam. (including lvs); outer surface of lvs with paper-thin grey tomentum

c. 500 spp., Eurasia, Africa, Australasia. Native spp. 9, naturalised 2.

The generic limits of Helichrysum are not clearly defined - in general Helichrysum differs from Cassinia in the absence of receptacular scales and from Gnaphalium in having fewer ♀ than ⚥ florets. The placement of many indigenous spp. is unsatisfactory and many "intergeneric" hybrids are recorded. The whole group requires further study particularly in relation to the complex in Australia. Most of the indigenous spp. are probably closest to the anaphaloid cudweeds now treated in Anaphalis.

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