Volume III (1980) - Flora of New Zealand Adventive Cyperaceous, Petalous & Spathaceous Monocotyledons
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Key to Genera

KEY TO GENERA

KEY TO GENERA

† Treated in Vol. II.     ζ See note below key.

Key

1
Perianth of separate calyx and corolla, or sepaloid, dry and glumaceous, or of minute scales or bristles, or 0
2
Perianth composed entirely of petaloid segments
60
2
Plants herbaceous (terrestrial or aquatic)
3
Plants arborescent or high-climbing lianes
55
3
Land plants, or aquatics firmly rooted in soil with floating leaves or stems, or leaves raised above the water; leaves if narrow, not translucent
4
Aquatic plants, free-floating on surface of fresh water or rooted in soil and normally entirely submersed or intertidal; leaves mostly filamentous to ribbon-like, usually ± translucent
44
4
Flowers inconspicuous or ± hidden, greenish or brownish
5
Flowers conspicuous, white or coloured, or minute but enclosed within a conspicuous spathe
33
5
Perianth-segments usually not obscured by bracts, though small
6
Perianth-segments 0, or microscopic and ± hidden
12
6
Fruit a capsule
7
Fruit is not a capsule
10
7
Flowers several to numerous, rarely solitary, usually < 5 mm long
8
Flowers solitary, usually >5 mm long
9
8
Capsule many-seeded; leaves (or basal sheaths) glabrous
Capsule 3-seeded; leaves sparsely hairy
9
Green leaves 2 or more on each shoot; floral bracts < tepals; seeds tailed
Green leaves 1–3 on each shoot; floral bracts = or > tepals; seeds without tails
10
Leaves elliptic to broadly ovate
Leaves of linear order
11
11
Flowers in compact, globose, obviously unisexual heads
Flowers bisexual in a narrow raceme
12
Leaves ± plano-convex in T. S., spongy within, or all reduced to sheaths spaced at intervals along stem
13
Leaves flat, terete or setaceous, or reduced to sheaths crowded at stem-base
16
13
Flowers and associated structures all thread-like, densely crowded in large cylindric velvety-brown inflorescences
Flowers and associated bracts and glumes not wholly thread-like; inflorescences of several to many ovate spikelets, not velvety
14
14
Stems unbranched
Stems branched
15
15
Inflorescence a panicle
SPORADANTHUS†
Inflorescence of distant spikelets towards stem-tip
EMPODISMA†ζ
16
Flowers in many-glumed spikelets
17
Flowers in a solitary, complex but superficially spike-like inflorescence enclosed within 2–3 glume-like floral bracts
32
17
Nut enclosed in an utricle; flowers all unisexual
18
Nut not enclosed in an utricle; flowers predominantly hermaphrodite
19
18
Rhachilla not produced beyond utricle; flowers usually in several spikes but occasionally in a solitary spike
Rhachilla produced beyond utricle into a hook; flowers in a solitary spike
UNCINIA†
19
Almost all glumes of spikelet flower-bearing
20
Only a few glumes in middle of upper part of spikelet flower-bearing
24
20
Glumes distichous; spikelets flattened
Glumes spirally arranged; spikelets terete
21
21
Leaves glabrous, or scabrid only towards tip, or reduced to basal sheaths
22
Leaves harshly scabrid throughout or pubescent
23
22
Inflorescence usually bracteate, terminal but often apparently lateral; nut without persistent style-base
Inflorescence ebracteate, terminal and appearing so; nut crowned by enlarged persistent style-base
23
Leaves harshly scabrid on margins and keel; style without basal hairs, continuous with nut
Leaves softly pubescent; style with a basal ring of drooping hairs, articulate on nut
24
Perianth 0, or of 6–3 non-plumose setae
25
Perianth of 6 plumose setae or 6 hypogynous scales
30
25
Leaves flattened, spirally arranged
26
Leaves terete, or flattened but distichous or reduced to basal sheaths
27
26
Stems terete; glumes spirally arranged
GAHNIA†
Stems 3-angled; glumes distichous
27
Rhachilla of spikelets zigzag
Rhachilla of spikelets straight
28
28
Persistent style-base ± ½ length of nut or shorter
Persistent style-base ± = body of nut in length
29
Leaves flattened
Leaves terete or reduced to basal sheaths
30
Perianth of plumose setae
Perianth of hypogynous scales
31
31
Leaves similar to stems, or reduced to basal sheaths
LEPIDOSPERMA†
Leaves unlike stems
32
Leaf-sheaths hyaline, pale green or colourless; glume-like floral bracts 2, appearing opposite
Leaf-sheaths opaque, light-brown; glume-like floral bracts (2)–3, markedly alternate
33
Perianth minute or 0; flowers aggregated on a spadix, ± enclosed in a large spathe
34
Perianth obvious; flowers not aggregated on a spadix though inflorescence occasionally enclosed at first by spathe-like bracts
38
34
Leaf-veins of same colour as lamina (excluding any spots); spathe usually white
Leaf-veins yellower or redder than lamina; spathe greenish to yellowish, or green and purple
35
35
Leaf-sheaths spotted
36
Leaf-sheaths not spotted
37
36
Leaves dissected; spathe green and purple
Leaves entire; spathe green to yellowish
37
Leaves peltate
Leaves not peltate
38
Flowers with large showy petaloid staminodia
39
Flowers without staminodia
40
39
Anthers 2-locular; sepals united in a tube
Anthers 1-locular; sepals free
40
Flowers in 2 divergent spikes; perianth of a single white "segment"
Flowers solitary, or in panicles or cymes; perianth of 3 green sepals and 3 white, pink or yellow petals
41
41
Carpels united in a capsule
42
Carpels free
43
42
Leaves cauline; land plant
Leaves basal; aquatic plant
43
Flowers white or pink, in panicles
Flowers yellow, solitary
44
Plants submersed, attached by roots, differentiated into stem and leaves or bracts
45
Plants floating free on fresh water, composed of oval green platelets budding from lateral pouches
53
45
Leaf-margins minutely serrate, laminae not crimped
46
Leaf-margins smooth, or serrate if laminae crimped
48
46
Leaves in whorls of 3–8, occasionally some opposite at base of stem
Leaves in whorls of 2–3, occasionally some opposite at base of stem; or leaves alternate
47
47
Leaves whorled, lax, not recurved; flowers obvious
Leaves alternate, stiff, recurved; flowers minute
48
Plants of sites below regular high tide mark, usually forming grass-like sward; leaves linear but not filiform, with numerous transverse veinlets at right angles to midrib and strong marginal nerves
Plants of brackish or fresh water; leaves usually distributed on slender stems, or, if closely crowded, almost filamentous, without numerous transverse veinlets and strong marginal nerves
49
49
Peduncles, long, thread-like, becoming spirally coiled as fruit matures; flowers 2, sessile and close together, each producing (2)–4–8–(16) obviously stalked achenes
Peduncles mostly short, never spirally coiled; flowers 1–many, each producing at most 4 sessile or obscurely stalked achenes
50
50
Leaves without sheaths or stipules; plant rarely as much as 5cm high
Leaves with stipules, sometimes sheathing; stem-internodes mostly easily visible, many >1 cm long
51
51
Flowers bisexual, tetramerous, many, in obvious, fully exserted, continuous or interrupted spikes
Flowers unisexual, not tetramerous, solitary or few in inconspicuous clusters
52
52
Leaves in false whorls of 3–4; stipules free
Leaves obviously alternate, mostly distant; stipules adnate to leaf-base
53
Roots absent; platelets deeply keeled
WOLFFIA†
Roots present; platelets flat to slightly biconvex, without deep keel
54
54
Roots 1 per platelet
LEMNA†
Roots 2 or more per platelet
55
Plant arborescent; leaves compound
56
Plant a high-climbing liane; leaves simple
59
56
Leaves palmate
57
Leaves pinnate
58
57
Petioles toothed or rough on margins
Petioles usually spiny on margins
58
Leaflets reduplicately inserted on axis (A-shaped in T.S.); trees monoecious; spathes subtending inflorescence2
RHOPALOSTYLIS†
Leaflets induplicately inserted on axis (V-shaped in T.S.); trees dioecious; spathe subtending inflorescence solitary
59
Stem twiner; leaves reticulately veined, margins smooth; inflorescence laxly racemose
Root climber; leaves parallel-veined, margins scabrid; inflorescence a dense spadix
60
Flowers actinomorphic or zygomorphic but without a labellum; stamens 6 or 3, usually free but filaments sometimes united
61
Flowers usually strongly zygomorphic with one petal differentiated as labellum; stamens 1–2 on a central bilaterally symmetric column
112
61
Ovary superior
62
Ovary inferior
88
62
Leaves all basal
63
Leaves all cauline, or basal and cauline, sometimes bract-or scale-like
79
63
Leaves equitant
PHORMIUM†
Leaves not equitant
64
64
Flowers white, blue, pink or purple, greenish-yellow or greenish-fawn to maroon
65
Flowers bright yellow, orange or red
76
65
Perianth-tube 0, or much < lobes
66
Perianth-tube long, much > lobes
73
66
Flowers in umbels, spikes, or solitary
67
Flowers in racemes or panicles
71
67
Plant with onion smell
68
Plant without onion smell
69
68
Perianth-segments free, or united only at base; flowers many, corona 0
Perianth-segments united to a short tube; flowers one, or many with corona
69
Stoloniferous aquatic; leaf-petioles swollen, bladder-like
Bulbous or rhizomatous land plants; leaf-petioles 0
70
70
Plants bulbous; stamens erect
Plant rhizomatous; stamens declinate
71
Perianth-lobes with a conspicuous green or purple median vein; plant with fleshly-fibrous roots
Perianth-lobes without median stripe or striped only on back; plant bulbous
72
72
Perianth-lobes white with green median dorsal stripe; staminal-filaments flattened
Perianth-lobes variously coloured, blue, rose or white, without green median stripe; staminal-filaments filiform or dilated only at base
73
Plant with bulbs or corms, summer-green; leaves flaccid
74
Plant tufted or with rhizomes, evergreen; leaves stiff
75
74
Flowers bright blue to violet-blue; perianth flask-shaped, lobes to 1/10 length of tube
Flowers light blue, pink, heliotrope, purple or white; perianth campanulate, lobes ¼ length of tube
75
Perianth-tube fleshly or almost absent
ASTELIA†
Perianth-tube deeply cup-shaped and membranous
76
Perianth-tube much > lobes; leaf-margins with pungent teeth
Perianth-tube 0, or much < lobes; leaf-margins smooth
77
77
Inflorescence branched, corymbose; flowers orange-red
Inflorescence usually a raceme or panicle; flowers yellow
78
78
Leaves strongly plicate; scape hairy
Leaves not plicate; scape glabrous
79
Cladodes needle-like or leaf-like, in axils of scale-like leaves
Cladodes absent, leaves well-developed
80
80
Leaf-petiole twisted to bring convex abaxial leaf-surface uppermost
Leaf-petiole straight or 0
81
81
Plants large, > 20 cm high; cauline leaves well-developed; flowers in panicles or racemes, or solitary and axillary or >3 cm diam.
82
Plants small, < 10 cm high; cauline leaves bract-like; flowers terminal, solitary, < 1 cm diam.
87
82
Plants bulbous; flowers > 5 cm long
Plants rhizomatous, or tuberous, or without underground storage organs, or arborescent; flowers < 5 cm long
83
83
Flowers in axillary clusters of 2–4; leaves linear elliptic
Flowers in racemes or panicles; leaves linear to lanceolate
84
84
Flowers red, in dense racemes; rocky places, outlying islands
Flowers white or greenish-white, in open racemes or panicles; coastal rocks, forest margins, scrub etc., mainland
85
85
Leaves ± fleshly or flaccid; fruit a capsule
ARTHROPODIUM†
Leaves tough; fruit a berry
86
86
Stem woody, sometimes forming long trunk; midrib not scabrid abaxially
Stem scarcely woody, almost entirely enclosed in leaves, midrib scabrid abaxially
DIANELLA†
87
Plant cormous; flowers distinctly pedicellate; segments to 5 mm long
Plant rhizomatous; flowers ± sessile; segments 10–15 mm long
88
Stamens 3; leaves mostly equitant
89
Stamens 6; leaves not equitant
105
89
Flowers solitary, often sessile within each pair of spathe-valves, not short-lived
90
Flowers 2 or more, pedicellate, within each pair of spathe-valves, ± fugitive, each pedicel subtended by a membranous bract
101
90
Inflorescence spicate, or a stiff panicle with flowers spicately arranged
91
Inflorescence a lax panicle with slender pendulous branches, or flowers arising singly from a corm
100
91
Style-branches simple (stigmas 3)
92
Style-branches bifid (stigmas 6)
99
92
Perianth-tube straight in upper wider part
93
Perianth-tube curved in upper wider part
98
93
Outer spathe-valve ovate, truncate, either shortly apiculate or notched or toothed but not lacerate at tip, glabrous, usually < 12 mm long
94
Outer spathe-valve lanceolate or cuspidate, entire or lacerate at tip, hairy or glabrous, usually 12–25 mm long
96
94
Flowers in a ± flattened panicle, main inflorescence-axis and each branch terminating in a zigzag portion; capsule 3–5-seeded
Flowers in a usually distichous or spirally arranged spike; capsule many-seeded
95
95
Perianth-lobes indistinctly nerved; stamens symmetrical
Perianth-lobes distinctly nerved; stamens asymmetric
96
Perianth-tube ± 2.5 cm long, very narrow-cylindrical throughout; plant rhizomatous
Perianth-tube < 1.5 cm long, funnel-shaped above; plant cormous
97
97
Leaves glabrous, flat
Leaves hairy, plicate
98
Perianth-tube dilating gradually; upper perianth-lobes not hooded; spathe-valves herbaceous, green
Perianth-tube narrow-cylindrical in lower half, abruptly dilated at middle; upper perianth-lobe ± hooded; spathe-valves membranous brownish
99
Flowering stems robust, 75–200 cm high; flowers many
Flowering stems slender, usually < 30 cm high; flowers 2–7
100
Flowers in a lax panicle, bell-shaped, on pendulous branches; large tussock
Flowers arising singly, 1- several from a corm, funnel-shaped, perianth-lobes ± spreading; low growing, ± inconspicuous
101
Basal leaf 1; plant cormous
Basal leaves many, often in flattened fans; plant rhizomatous, or annual
102
102
Flowers large and showy; perianth-lobes in 2 very dissimilar series; style-branches petaloid
Flowers comparatively small; perianth-lobes in 2 similar series, outer occasionally differing in size from inner; style-branches not petaloid
103
103
Flowers white; outer perianth-lobes shorter and narrower than inner
LIBERTIA†
Flowers creamy-yellow often purple-striped, violet or blue; perianth-lobes equal in length
104
104
Style with short style-branches; flowers creamy-yellow with purple stripes or light violet-blue; base of leaf-fan light green
Style unbranched with 3 lobes at tip; flowers deep blue and base of leaf-fan bright reddish-purple
105
Plant bulbous or cormous
106
Plant with tubers and/or rhizomes, or without any subterranean storage organs
109
106
Perianth-lobes pink, or white tipped with green
107
Perianth-lobes yellow to pure white
108
107
Scape solid; flowers c. 10 cm long, pink
Scape hollow; flowers c. 2 cm long, white with green-tipped perianth-lobes
108
Flowers with corona; plant bulbous
Flowers without corona; plant cormous
109
Leaves stiff and ± fleshly or leathery, margins often toothed, tip horny and pungent; flowers greenish, brownish or whitish
110
Leaves soft, not toothed or pungent-tipped; flowers red or orange
111
110
Inflorescence with bulbils, or without bulbils and leaf-margins sharply but finely serrate
Inflorescence without bulbils; leaf-margins with prominent horny teeth
111
Plant with chlorophyll, tuberous and/or rhizomatous
ALSTROMERIA
Plant without chlorophyll and without subterranean storage organs
112
Plant terrestrial, sometimes on rotting logs, etc.; stems annual
113
Plant epiphytic or rupestral, sometimes persisting after falling to ground; stems perennial
131
113
Leaves with chlorophyll
114
Leaves without chlorophyll, sometimes reduced to pale scales
129
114
Green leaf on flower-stem solitary; cauline bracts occasionally present
115
Green leaves on flower-stem >1 (sometimes only 1 in Lyperanthus)
123
115
Leaf flat or channelled
116
Leaf tubular
122
116
Leaf short and broad, lamina-length mostly < 2 × breadth
117
Leaf relatively long and narrow
119
117
Peduncle shorter than flower, lengthening in fruit; flower usually solitary
CORYBAS†
Peduncle longer than flower; flowers 1-several
118
118
Flower usually solitary; labellum with many, ± clavate calli in longitudinal rows
ADENOCHILUS†
Flowers usually > 1; labellum without such rows of calli
ACIANTHUS†
119
Leaves and/or stem minutely hairy
CALADENIA†
Leaves and stem glabrous
120
120
Flower almost regular as to perianth; labellum little differentiated
THELYMITRA†
Flower with well differentiated labellum
121
121
Labellum pendent from flower with conspicuous reddish beard
CALOCHILUS†
Labellum uppermost, its peltate tuberculate lamina attached by slender claw
CALEANA†
122
Dorsal sepal hooded; labellum below column
MICROTIS†
Dorsal sepal not hooded; labellum above column
PRASOPHYLLUM†
123
Green leaves 2, mostly rather broad; stigma adjacent to anther; flowers solitary or 1–3
124
Green leaves usually > 2, ± linear, or if broad then either flowers many or stigma distant from anther
126
124
Leaves glandular-hairy; petals and sepals mostly white
APOROSTYLIS†
Leaves glabrous; flowers greenish
125
125
Leaves close together; peduncle < leaves at flower, lengthening in fruit rarely > 1 flower
CHILOGLOTTIS†
Leaves well spaced; peduncle much > leaves at flower, sometimes with leaf-like bract; often > 1 flower
LYPERANTHUS†
126
Flower solitary, or few in short inflorescence
PTEROSTYLIS†
Flowers several to many in long inflorescence
127
127
Flowers always inverted so that labellum lies above column
CRYPTOSTYLIS†
Flowers not inverted on the scape; labellum always below column
128
128
Perianth-segments unequal; lateral sepals filiform, 1.5–3 cm long
ORTHOCERAS†
Perianth-segments subequal, rarely > 6 mm long
SPIRANTHES†
129
Flower solitary, barely emerging from litter
CORYBAS†
Flowers several to many, on erect stem
130
130
Sepals and lateral petals fused to form tube; rhizome 1–5 cm diam.
GASTRODIA†
Sepals and lateral petals free; rhizome rarely > 5 mm diam.
YOANIA†
131
Leaves mostly < 2.5 cm long, each attached to pseudobulb
BULBOPHYLLUM†
Leaves mostly > 2.5 cm long, without pseudobulbs
132
132
Stem very short, with crowded, ± oblong, distichous leaves
DRYMOANTHUS†
Stem long, with well spaced, narrow-linear leaves
133
133
Stems freely branched; flowers few together
Stems usually unbranched; flowers many together
EARINA†

ζEmpodisma Johnson et Cutler (Kew Bull. 28, 1973, 383) was segregated from Calorophus Labill., which is now considered to be restricted to Tasmania. They summarised in a table the many differences in floral morphology and culm anatomy between the two genera. One sp. of Empodisma is endemic to S. W. Australia, the other, E. minus (Hook. f.) Johnson et Cutler based on Calorophus minor Hook. f., is widespread in S. E. Australia, Tasmania and N.Z.

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