Volume IV (1988) - Flora of New Zealand Naturalised Pteridophytes, Gymnosperms, Dicotyledons
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Myosotis L.

MYOSOTIS L.

Annual, biennial or perennial, ± hairy herbs. Habit from cushion form to erect or sprawling. Infl. usually cymose, ebracteate or bracteate, often enlarged at fruiting. Calyx 5-toothed, often accrescent. Corolla ± rotate; tube short, infolded to form 5 scales at throat; limb of 5 equal lobes, patent or concave, contorted in bud, blue, purple, white, yellow, brown or orange. Stamens with anthers either partly below scales (sect. Myosotis) or wholly above them (sect. Exarrhena). Ovary 4-celled; style slender. Nutlets 4, ovoid, brown or black, smooth and glossy, often with a distinct rim.

Key

1
Anthers included, partially or wholly below corolla scales
2
Anthers exserted, wholly above corolla scales
24
2
Cymes ebracteate or with bracts at base only, elongating greatly after flowering; corolla blue or becoming blue (occasional white-flowered mutants in a population)
3
Cymes bracteate or if ebracteate, then not elongating greatly after flowering; corolla usually white, yellow, or brown to almost black, or rarely blue
8
3
Plants mostly of wet places; calyx with straight appressed hairs
4
Plants not of wet places; calyx with spreading hairs, some hooked or curled
5
4
Calyx teeth c. ⅓ length of calyx at anthesis; corolla limb 5-10 mm diam.
Calyx teeth c. 1/2 length of calyx at anthesis; corolla limb 2-5 mm diam.
5
Corolla limb 6-11 mm diam., spreading or flat
Corolla limb < 5 mm diam., concave
6
6
Fruiting cymes < to slightly > leafy part of stem; fruiting pedicels > calyx (at least in lower part of infructescence)
Fruiting cymes usually > leafy part of stem; fruiting pedicels < calyx
7
7
Veins of lower lf surface with some hooked hairs; corolla always blue
Veins of lower lf surface without hooked hairs; corolla firstly cream or yellow, then becoming blue
8
Fls either in bracteate cymes on decumbent or lax branches, or solitary and terminal
9
Fls in ebracteate cymes on erect or ascending branches
18
9
Fls solitary, terminal; lvs often tightly imbricate; twigs cylindric
10
Infl. of several fls with bracts; lvs not tightly imbricate; twigs not conspicuously cylindric
12
10
Lvs sparsely hairy, soon glabrous; only lower extremity of anthers at scale-level
Lvs densely clothed in persistent hairs; only upper tip of anthers above scale-level
11
11
Lvs narrow-elliptic, pointed; fls yellow fading to white, often both colours present on a cushion
Lvs broadly spathulate, often very broadly so, obtuse to rounded; all fls white
12
Corolla 4-8 mm diam.; lobes flat, broad; anthers > 1 mm long, tips protruding from tube; style much > calyx
13
Corolla 1-4 mm diam.; lobes ± concave, narrow; anthers < 1 mm long, wholly included; style slightly > or = calyx
15
13
Hairs soft, spreading; lvs obovate
Hairs stiff, appressed; lvs elliptic or lanceolate
14
14
Lateral branches extending well beyond central rosette
Lateral branches short and mostly included in leafy cushion
15
Calyx lobed almost to base; lobes widespreading at fruiting; calyx hairs appressed, uniformly short, stiff, forming sparse but even cover
Calyx lobed to c. 1/2 way; lobes ± erect at fruiting; calyx hairs ± spreading, uniform or not, ± aggregated in longitudinal strips
16
16
Fls usually white to very pale yellow; hairs not crowded, not very fine and silky
A
Fls 0.5-1 mm diam.
var. minutiflora
Fls > 1 mm diam.
B
B
Calyx hairs few, stiff and short
var. glauca
Calyx hairs many, both short and long
C
C
Hairs with soft, ± flexuous tip; nutlets quite immersed in long calyx
var. drucei
Hairs strigose; calyx tube short, nutlets protruding
var. pygmaea
Fls usually blue; hairs crowded except on lower surface of mature lvs, very fine and silky
17
17
Rosette lvs usually > 3 cm long; cymes ebracteate except at base
Rosette lvs usually < 2 cm long; cymes bracteate to near apex
18
Calyx hairs of 2 sharply distinct kinds: (a) short, fine, straight, closely appressed; (b) long, coarse, hooked, spreading
A
Stem lvs without hooked hairs on undersurface
var traversii
Stem lvs with many, long, hooked hairs on undersurface
B
B
Lf hairs long and spreading, those on undersurface of rosette lvs retrorse; infl. usually capitate even at fruiting
var cantabrica
Lf hairs short and smoothly appressed, not retrorse on undersurface of rosette lvs; infl. usually elongate at fruiting
var cinerascens
Calyx hairs of a continuous range of sizes, many or all of the longer ones not hooked
19
19
Hairs on undersurface of rosette lvs regularly retrorse
20
Hairs on undersurface of rosette lvs not regularly retrorse
22
20
Calyx 3-6 mm long, deeply lobed, with many hairs definitely hooked; anther tips not protruding above scales
Calyx 5-9 mm long, lobed to about 1/2 way, with few if any hairs definitely hooked; anther tips projecting above scales
21
21
Hairs crowded; corolla c. 7 mm diam.
Hairs sparse; corolla c. 10 mm. diam.
22
Hairs sparse; infls long; internodes between frs > calyx
Hairs crowded; infls compact; only lowest if any internodes > calyx
23
23
Hairs overlapping and hiding epidermis on shoots and lvs; whole plant white
Hairs overlapping but not hiding epidermis on shoots and lvs; whole plant ashen grey
24
Cymes wholly, or occasionally partly, ebracteate
25
Each fl. associated with a bract-like lf
39
25
Infl. usually of < 8 narrow-tubed fls, with up to 3 of lowest fls bracteate; lateral branches ± decumbent, rarely > 8 cm long; hairs flexuous, neither hooked nor retrorse, sparse or 0 on undersurface of rosette lvs
A
Plant compact; hairs ± overlapping; calyx lobed to ⅓
var lyallii
Plant lax; hairs few and widely spaced; calyx lobed to at least 1/2
var townsonii
Plant not with all characters as above in combination
26
26
Infl. lax at fruiting, with most internodes then > calyx
27
Infl. dense at fruiting, with most internodes then < calyx
31
27
Calyx lobed to about 1/2; calyx lobes broad, ribbed; lf hairs flexuous, none retrorse
Calyx lobed almost to base; calyx lobes narrow, without ribs; lf hairs straight, or if flexuous then some retrorse
28
28
Hairs on undersurface of rosette lvs retrorse
29
Hairs on undersurface of rosette lvs spreading or antrorse
30
29
Corolla nearly campanulate; corolla tube > calyx
Corolla rotate; corolla tube < calyx
30
Lvs ± elliptic, very gradually narrowed to petiole; calyx hairs long, flexuous to slightly hooked, crowded
Lvs orbicular to broad-elliptic, narrowed abruptly to petiole; calyx hairs short, stiff, sparse
A
A Corolla tube ± cylindric, much longer than wide, with scales far above calyx tips
var. pottsiana
Corolla tube wide-funnelform, barely as long as wide, with scales at or below calyx tips
B
B
B Corolla 6-8 mm diam., with scales c. at level of calyx tips; stamens exceeding corolla
var. petiolata
Corolla 10-15 mm diam., with scales much below calyx tips; stamens not exceeding corolla
var. pansa
31
Filaments much > anthers
32
Filaments c. = or slightly > anthers
35
32
Hairs all short, stiff, closely appressed, sparse
33
Hairs mostly flexuous or spreading, with some appressed and crowded
34
33
Fls white; corolla tube wide-funnelform; calyx hairs not hooked
Fls yellow; corolla tube cylindric; calyx with some hooked hairs
34
Lf hairs appressed; fls yellow; filaments standing well above corolla
Lf hairs spreading; fls white; filaments reaching to level of corolla lobes
35
Rosette lvs broader than stem lvs; retrorse hairs present
36
Rosette lvs narrow like stem lvs; retrorse hairs 0
38
36
Hairs stiff, smoothly appressed
Hairs ± flexuous, not smoothly appressed
37
37
Infl. usually much-branched; corolla white; corolla tube < calyx; calyx 4-5 mm long at fruiting
Infl. simple or once-forked; corolla yellow to brownish orange to brown to deep purple; corolla tube much > calyx; calyx 6-9 mm long at fruiting
38
Hairs overlapping and hiding epidermis; whole plant silvery
Hairs barely overlapping, not hiding epidermis; surfaces grey-green
39
Plant erect or sprawling, without roots among stems lvs; stem hairs short and closely appressed; calyx hairs uniformly short and straight; corolla broadly open; stigma capitate
Plant ± creeping, with adventitious roots among stem lvs; stem hairs slightly spreading to shaggy; calyx hairs not uniform in size, the largest ± hooked; corolla narrow; stigma clavate
A
A Plant straggling, hispid, especially on stems; pedicel much > calyx at fruiting; corolla c. 3 mm diam.
var spathulata
Plant creeping, rooting freely, hairs shorter; pedicel c. = calyx at fruiting; corolla up to c. 2 mm diam.
var radicata

c. 50 spp., temperate Eurasia, tropical montane Africa, South Africa, Australasia. Native spp. 34, naturalised 6.

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