Volume IV (1988) - Flora of New Zealand Naturalised Pteridophytes, Gymnosperms, Dicotyledons
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Myrtaceae

MYRTACEAE

Evergreen shrubs or trees. Lvs simple, usually entire, sometimes finely toothed in juvenile and intermediate lvs, exstipulate, very rarely stipulate, generally coriaceous, opposite, or alternate (often in adult shoots), often dotted with pellucid glands. Fls solitary in cymes, often in panicles, actinomorphic, usually ⚥. Hypanthium (calyx tube) ± adnate to ovary, the lower part sometimes narrow and forming a pseudopedicel; lobes (2)-4-5-(many), sometimes calyptrate, sometimes calyx rim-like. Petals as many as calyx lobes or rarely 0, imbricate, sometimes caducous, sometimes calyptrate. Stamens usually numerous; filaments free or connate at base, sometimes in bundles opposite petals; anthers small, usually dehiscing by longitudinal slits, rarely by apical pores; connective gland-tipped. Ovary inferior; ovules usually numerous, rarely few or 1, mostly with axile, rarely parietal, placentation; style simple; stigma capitate. Fr. a loculicidal capsule or berry-like. Seeds usually numerous; endosperm 0 or scanty.

SYNOPSIS

is represented in N.Z. by , the classical myrtle of the Mediterranean region. Although not wild, it long persists as a relic of cultivation and sometimes regenerates around mature plants. It is distinguished from all other spp. of its subfamily in N.Z. by the following combination of characters: a sweetly aromatic shrub; lvs ovate, acute; fls axillary, single or paired; petals almost patent, prominent, white; frs c. 1 × 0.7 cm, oblong-ellipsoid, black, with prominently accrescent calyx lobes.

Key

1
Fr. a capsule, coriaceous to woody
2
Fr. fleshy and berry-like
10
2
Fls with petals usually insignificant and much < the conspicuous stamens
3
Fls with conspicuous petals > to slightly < stamens
7
3
Calyx forming a cap or operculum shed transversely at anthesis
Calyx without a cap, the sepals either persistent or shed separately
4
4
Lvs opposite; fls in terminal cymes, sometimes in corymbs, or racemes, not contiguous
5
Lvs alternate; fls contiguous in spikes or in elongated or rounded heads
6
5
Hypanthium smooth; shrubs, or if trees then fls nearly always red, extremely rarely white and then bark soft, spongy and whitish
Hypanthium ribbed; trees with white fls; bark smooth or rough, flaky, fibrous, or hard, pink or shades of grey
6
Lvs usually > 4 cm long; calyx lobes persistent to fruiting; capsule woody, long-persistent
Lvs usually < 4 cm long; calyx lobes usually deciduous; capsule coriaceous
7
Ovary 1-celled
Ovary 5- or more celled
8
8
Fls few to rather numerous, in dense globular heads; ovary 3-celled, each cell with 2-6 erect ovules on a basal placenta
Fls solitary or few in clusters, if few then not in globular heads; ovary usually 5-celled, each cell with many pendulous ovules on an apical placenta
9
9
Filaments in bud incurved towards stigma; capsule usually woody, rarely coriaceous
Filaments in bud inflexed to bring anthers below stigma; capsule coriaceous
10
Fls axillary, usually solitary but sometimes 2 -3 in cymes
11
Fls in terminal cymes or panicles
14
11
Fls usually > 15 mm diam.; fr. > 20 mm long
Fls < c. 12 mm diam.; fr. < 15 mm long (at least in N.Z.)
12
12
Corolla campanulate with strongly concave, imbricate petals; filaments flattened or dilated
Corolla opening widely, with petals ± patent and scarcely imbricate; filaments terete and filiform
13
13
Sepals and petals 4; ovules in 1 series
LOPHOMYRTUS†
Sepals and petals 5; ovules in 2 series
NEOMYRTUS†
14
Calyx lobes connate into a calyptrum; anther sacs divergent
Calyx lobes free, often rudimentary; anther sacs parallel

c. 100 genera, 3000 spp., mainly S. Hemisphere and tropical.

The spp. which are recorded here as wild are usually not conspicuously and widely naturalised. Genera listed for the 2 subfamilies below include a number which, although conspicuous in cultivation and as relics of cultivation in old gardens, have no wild members.

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